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Cortical Alveolum
The cortical alveolum () is a cellular organelle consisting of a vesicle located under the cytoplasmic membrane, to which they give support. The term " corticate" comes from an evolutionary hypothesis about the common origin of kingdoms Plantae and Chromista, because both kingdoms have cortical alveoli in at least one phylum. At least three protist lineages exhibit these structures: Telonemia, Alveolata and Glaucophyta. Definition Cortical alveoli have been defined as flattened membranous sacs or vesicles that strengthen the cellular cortex through the firm fixation to the underlying membrane and microtubules. They typically form a continuous layer that acts as a flexible film, although they can also constitute a semi-rigid structure or the scales of a theca. Occurrence Cortical alveoli are the defining cellular characteristic of the protist clade Alveolata, otherwise united solely by molecular phylogeny. The name 'Alveolata' is a reference to this trait, present in all re ...
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EM Alveoli Ciliate Paramecium Putrinum
EM, Em or em may refer to: Arts and entertainment Music * Em, the E minor musical scale * Em, the E minor chord * Electronic music, music that employs electronic musical instruments and electronic music technology in its production * Encyclopedia Metallum, an online metal music database * Eminem, American rapper Other uses in arts and entertainment * ''Em'' (comic strip), a comic strip by Maria Smedstad Companies and organizations * Em (restaurant), a restaurant in Mexico City * Aero Benin (IATA code), a defunct airline * Empire Airlines (IATA code), a charter and cargo airline based in Idaho, US * Erasmus Mundus, an international student-exchange program * ''Estado de Minas'', a Brazilian newspaper * European Movement, an international lobbying association * ExxonMobil, a large oil company formed from the merger of Exxon and Mobil in 1999 * La République En Marche! (sometimes shortened to "En Marche!"), a major French political party Economics * Emerging markets, nations und ...
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Molecular Phylogeny
Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to determine the processes by which diversity among species has been achieved. The result of a molecular phylogenetic analysis is expressed in a phylogenetic tree. Molecular phylogenetics is one aspect of molecular systematics, a broader term that also includes the use of molecular data in taxonomy and biogeography. Molecular phylogenetics and molecular evolution correlate. Molecular evolution is the process of selective changes (mutations) at a molecular level (genes, proteins, etc.) throughout various branches in the tree of life (evolution). Molecular phylogenetics makes inferences of the evolutionary relationships that arise due to molecular evolution and results in the construction of a phylogenetic tree. History The theoretical framew ...
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Stramenopiles
The stramenopiles, also called heterokonts, are Protist, protists distinguished by the presence of stiff tripartite external hairs. In most species, the hairs are attached to flagella, in some they are attached to other areas of the cellular surface, and in some they have been secondarily lost (in which case relatedness to stramenopile ancestors is evident from other shared cytological features or from genetic similarity). Stramenopiles represent one of the three major clades in the SAR supergroup, SAR Supergroup (biology), supergroup, along with Alveolate, Alveolata and Rhizaria. Stramenopiles are Eukaryote, eukaryotes; most are single-celled, but some are multicellular including some large seaweeds, the brown algae. The group includes a variety of algal protists, heterotrophic flagellates, opalines and closely related proteromonad flagellates (all Endosymbiont, endobionts in other organisms); the actinophryid Heliozoa, and oomycetes. The tripartite hairs characteristic of the ...
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Supergroup (biology)
A supergroup or super-group, in systematics, is a large group of organisms that monophyletic, share one common ancestor and have important defining characteristics. It is an informal, mostly arbitrary rank in biology, biological Taxonomy (biology), taxonomy that is often greater than phylum or kingdom (biology), kingdom, although some supergroups are also treated as phylum, phyla. Eukaryotic supergroups Since the decade of the 2000s, the Eukaryote#Phylogeny, eukaryotic tree of life (abbreviated as eToL) has been divided into 5–8 major groupings called 'supergroups'. These groupings were established after the idea that only monophyletic groups should be accepted as ranks, as an alternative to the use of paraphyletic kingdom Protista. In the early days of the eToL six traditional supergroups were considered: Amoebozoa, Opisthokonta, "Excavata", Archaeplastida, "Chromalveolata" and Rhizaria. Since then, the eToL has been rearranged profoundly, and most of these groups were found as pa ...
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TSAR
Tsar (; also spelled ''czar'', ''tzar'', or ''csar''; ; ; sr-Cyrl-Latn, цар, car) is a title historically used by Slavic monarchs. The term is derived from the Latin word '' caesar'', which was intended to mean ''emperor'' in the European medieval sense of the term—a ruler with the same rank as a Roman emperor, holding it by the approval of another emperor or a supreme ecclesiastical official—but was usually considered by Western Europeans to be equivalent to "king". Tsar and its variants were the official titles in the First Bulgarian Empire (681–1018), Second Bulgarian Empire (1185–1396), the Kingdom of Bulgaria (1908–1946), the Serbian Empire (1346–1371), and the Tsardom of Russia (1547–1721). The first ruler to adopt the title ''tsar'' was Simeon I of Bulgaria. Simeon II, the last tsar of Bulgaria, is the last person to have held this title. Meaning in Slavic languages The title tsar is derived from the Latin title for the Roman emperors, ''c ...
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Phylum
In biology, a phylum (; : phyla) is a level of classification, or taxonomic rank, that is below Kingdom (biology), kingdom and above Class (biology), class. Traditionally, in botany the term division (taxonomy), division has been used instead of phylum, although the International Code of Nomenclature for algae, fungi, and plants accepts the terms as equivalent. Depending on definitions, the animal kingdom Animalia contains about 31 phyla, the plant kingdom Plantae contains about 14 phyla, and the fungus kingdom Fungi contains about eight phyla. Current research in phylogenetics is uncovering the relationships among phyla within larger clades like Ecdysozoa and Embryophyta. General description The term phylum was coined in 1866 by Ernst Haeckel from the Greek (, "race, stock"), related to (, "tribe, clan"). Haeckel noted that species constantly evolved into new species that seemed to retain few consistent features among themselves and therefore few features that distinguishe ...
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Colponemid
Colponemids are free-living alveolates, unicellular flagellates related to dinoflagellates, apicomplexans and ciliates. They are predators of other small eukaryotes, found in freshwater, marine and soil environments. They do not form a solid clade, but a sparse group of deep-branching alveolate lineages. Biology Colponemids are flagellated unicellular protists, 8 to 30 μm long, with rounded cell bodies that bear two heterodynamic flagella for rapid swimming. They are only present as swimming cells during their biological life cycle, life cycle. sexual reproduction, Reproduction or resting microbial cyst, cysts are unknown. Cell structure As other alveolates, all colponemids exhibit cortical alveoli and tubular crista, mitochondrial cristae. They present a three-membrane alveolar pellicle without fibrils, theca or micropores. Within the cell is a vesicular cell nucleus, nucleus with a central nucleolus, as well as a very large digestive vacuole (or food vacuole) located at ...
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Flagellate
A flagellate is a cell or organism with one or more whip-like appendages called flagella. The word ''flagellate'' also describes a particular construction (or level of organization) characteristic of many prokaryotes and eukaryotes and their means of motion. The term presently does not imply any specific relationship or classification of the organisms that possess flagella. However, several derivations of the term "flagellate" (such as " dinoflagellate" and " choanoflagellate") are more formally characterized. Form and behavior Flagella in eukaryotes are supported by microtubules in a characteristic arrangement, with nine fused pairs surrounding two central singlets. These arise from a basal body. In some flagellates, flagella direct food into a cytostome or mouth, where food is ingested. Flagella role in classifying eukaryotes. Among protoctists and microscopic animals, a flagellate is an organism with one or more flagella. Some cells in other animals may be flage ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Dinoflagellates
The Dinoflagellates (), also called Dinophytes, are a monophyletic group of single-celled eukaryotes constituting the phylum Dinoflagellata and are usually considered protists. Dinoflagellates are mostly marine plankton, but they are also common in freshwater habitats. Their populations vary with sea surface temperature, salinity, and depth. Many dinoflagellates are photosynthetic, but a large fraction of these are in fact mixotrophic, combining photosynthesis with ingestion of prey ( phagotrophy and myzocytosis). In terms of number of species, dinoflagellates are one of the largest groups of marine eukaryotes, although substantially smaller than diatoms. Some species are endosymbionts of marine animals and play an important part in the biology of coral reefs. Other dinoflagellates are unpigmented predators on other protozoa, and a few forms are parasitic (for example, '' Oodinium'' and '' Pfiesteria''). Some dinoflagellates produce resting stages, called dinoflagellate cysts ...
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Chromerid
Chrompodellids are a phylum of single-celled protists belonging to the Alveolata Supergroup (biology), supergroup. It comprises two different polyphyletic groups of flagellates: the colpodellids, phagotrophic predators, and the chromerids, photosynthetic algae that live as symbionts of corals. These groups were independently discovered and described, but molecular phylogenetic analyses demonstrated that they are intermingled in a clade that is the closest relative to Apicomplexa, and they became collectively known as chrompodellids. Due to the history of their research, they are variously known in biological classification as Chromerida or Colpodellida (International Code of Zoological Nomenclature, ICZN)/Colpodellales (International Code of Nomenclature for algae, fungi, and plants, ICN). Description and life cycle Chrompodellids are a phylum of unicellular Protist, protists containing two functionally different groups: the Photosynthesis, photosynthetic "chromerids" and the ...
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Colpodellid
Chrompodellids are a phylum of single-celled protists belonging to the Alveolata supergroup. It comprises two different polyphyletic groups of flagellates: the colpodellids, phagotrophic predators, and the chromerids, photosynthetic algae that live as symbionts of corals. These groups were independently discovered and described, but molecular phylogenetic analyses demonstrated that they are intermingled in a clade that is the closest relative to Apicomplexa, and they became collectively known as chrompodellids. Due to the history of their research, they are variously known in biological classification as Chromerida or Colpodellida (ICZN)/Colpodellales ( ICN). Description and life cycle Chrompodellids are a phylum of unicellular protists containing two functionally different groups: the photosynthetic "chromerids" and the predatory phagotrophic "colpodellids". Like other Alveolata, they present tubular mitochondrial cristae and highly flattened cortical alveoli with micro ...
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