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Chainosauria
Chainosauria is a large and speciose clade of anomodont therapsid that includes the highly diverse dicynodonts and a small number of closely related basal genera (to the exclusion of Venyukovioidea and more basal anomodonts)—although the total composition and taxonomic scope of Chainosauria is in flux. Chainosauria was named in 1923 to group together the dicynodonts and their close relatives, namely three small anomodont genera from South Africa (''Galechirus'', ''Galeops'', and ''Galepus'') that made up the now defunct group 'Dromasauria'. The name soon fell into disuse, however, as it was functionally replaced by Anomodontia. Chainosauria was later revived cladistically in 2009, preserving the association of dicynodonts and the 'dromasaurs' and has since served in effect as both a cladistic and a biogeographic counterpart to the Laurasian venyukovioids, with early chainosaurs appearing to have been a Gondwanan radiation. Other basal anomodonts, the anomocephaloids and ''Patra ...
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Venyukovioidea
Venyukovioidea is an infraorder of anomodont therapsids related to dicynodonts from the Permian of Russia. They have also known as Venjukovioidea, as well as by the similar names Venyukoviamorpha or Venjukoviamorpha in literature. This in part owes to a misspelling by Russian palaeontologist Ivan Efremov in 1940 when he mistakenly spelt ''Venyukovia'', the namesake of the group, with a 'j' instead of a 'y' (i.e. ''Venjukovia''), which permeated through subsequent therapsid literature before the mistake was caught and corrected. The order Ulemicia has also been coined for a similar taxonomic concept in Russian scientific literature, which notably excludes ''Suminia'' and ''Parasuminia''. Venyukovioidea includes the genera ''Venyukovia'', '' Otsheria'', '' Ulemica'', ''Suminia'' and ''Parasuminia'', all from Western Siberia. Historically, some of these genera have been placed in various families and subfamilies, including the Venyukoviidae/Venjukoviidae, Otsheriidae, and Ulemicii ...
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Anomodont
Anomodontia is an extinct group of non-mammalian therapsids from the Permian and Triassic periods. By far the most speciose group are the dicynodonts, a clade of beaked, tusked herbivores.Chinsamy-Turan, A. (2011) ''Forerunners of Mammals: Radiation - Histology - Biology''p.39.Indiana University Press, . Retrieved May 2012 Anomodonts were very diverse during the Middle Permian, including primitive forms like '' Anomocephalus'' and '' Patranomodon'' and groups like Venyukovioidea and Dromasauria. Dicynodonts became the most successful and abundant of all herbivores in the Late Permian, filling ecological niches ranging from large browsers down to small burrowers. Few dicynodont families survived the Permian–Triassic extinction event, but one lineage ( Kannemeyeriiformes) evolved into large, stocky forms that became dominant terrestrial herbivores right until the Late Triassic, when changing conditions caused them to decline, finally going extinct during the Triassic- ...
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Dicynodont
Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typically toothless beak, unique amongst all synapsids. Dicynodonts first appeared in Southern Pangaea during the mid-Permian, ca. 270–260 million years ago, and became globally distributed and the dominant herbivorous animals in the Late Permian, ca. 260–252 Mya. They were devastated by the end-Permian Extinction that wiped out most other therapsids ca. 252 Mya. They rebounded during the Triassic but died out towards the end of that period. They were the most successful and diverse of the non-mammalian therapsids, with over 70 genera known, varying from rat-sized burrowers to elephant-sized browsers. Characteristics The dicynodont skull is highly specialised, light but strong, with the synapsid temporal openings at the rear of the sku ...
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Dicynodontia
Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typically toothless beak, unique amongst all synapsids. Dicynodonts first appeared in Southern Pangaea during the mid-Permian, ca. 270–260 million years ago, and became globally distributed and the dominant herbivorous animals in the Late Permian, ca. 260–252 Mya. They were devastated by the end-Permian Extinction that wiped out most other therapsids ca. 252 Mya. They rebounded during the Triassic but died out towards the end of that period. They were the most successful and diverse of the non-mammalian therapsids, with over 70 genera known, varying from rat-sized burrowers to elephant-sized browsers. Characteristics The dicynodont skull is highly specialised, light but strong, with the synapsid temporal openings at the rear of the skul ...
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Middle Permian
The Guadalupian is the second and middle series/ epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian was previously known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by ...
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Cladistically
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived characteristics (synapomorphies'')'' that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms ''worms'' or ''fishes'' were used within a ''strict'' cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically, outside of cladistics, e.g. as a ' grade', which are fruitless to precisely delineate, especially when including extinct species. R ...
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Laurasian
Laurasia () was the more northern of two large landmasses that formed part of the Pangaea supercontinent from around (Mya), the other being Gondwana. It separated from Gondwana (beginning in the late Triassic period) during the breakup of Pangaea, drifting farther north after the split and finally broke apart with the opening of the North Atlantic Ocean c. 56 Mya. The name is a portmanteau of Laurentia and Asia. Laurentia, Avalonia, Baltica, and a series of smaller terranes, collided in the Caledonian orogeny c. 400 Ma to form Laurussia (also known as Euramerica, or the Old Red Sandstone Continent). Laurussia then collided with Gondwana to form Pangaea. Kazakhstania and Siberia were then added to Pangaea 290–300 Ma to form Laurasia. Laurasia finally became an independent continental mass when Pangaea broke up into Gondwana and Laurasia. Terminology and origin of the concept Laurentia, the Palaeozoic core of North America and continental fragments that now ...
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Gondwanan
Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages of break-up, involving the separation of Antarctica from South America (forming the Drake Passage) and Australia, occurred during the Paleogene. Gondwana was not considered a supercontinent by the earliest definition, since the landmasses of Baltica, Laurentia, and Siberia were separated from it. To differentiate it from the Indian region of the same name (see ), it is also commonly called Gondwanaland. Gondwana was formed by the accretion of several cratons. Eventually, Gondwana became the largest piece of continental crust of the Palaeozoic Era, covering an area of about , about one-fifth of the Earth's surface. During the Carboniferous Period, it merged with Laurasia to form a larger supercontinent called Pangaea. Gondwana (and Panga ...
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Autapomorphies
In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to the focal taxon (which may be a species, family or in general any clade). It can therefore be considered an apomorphy in relation to a single taxon. The word ''autapomorphy'', first introduced in 1950 by German entomologist Willi Hennig, is derived from the Greek words αὐτός, ''autos'' "self"; ἀπό, ''apo'' "away from"; and μορφή, ''morphḗ'' = "shape". Discussion Because autapomorphies are only present in a single taxon, they do not convey information about relationship. Therefore, autapomorphies are not useful to infer phylogenetic relationships. However, autapomorphy, like synapomorphy and plesiomorphy is a relative concept depending on the taxon in question. An autapomorphy at a given level may well be a synapomorphy at ...
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Anomocephaloids
Anomocephaloidea is a clade (evolutionary grouping) of anomodont therapsids that existed in Gondwana during the Middle Permian and includes two species, ''Anomocephalus africanus'' from South Africa and ''Tiarajudens eccentricus'' from Brazil, both of which are characterized by large body size and teeth that fit tightly together or occlude. Anomocephaloidea is among the most basal groups of anomodonts, the other being Venyukovioidea, which differs in being a Laurasian clade of mostly small-bodied species. Anomocephaloidea was named in 2011 with the discovery of ''Tiarajudens''; ''Anomocephalus'' had been known since 1999, but was unique among anomodonts until ''Tiarajudens'' was described. Both ''Anomocephalus'' and ''Tiarajudens'' were herbivores, although the latter possessed a pair of saber-like canine teeth that may have been used in display or combat with other individuals of the same species. Although Anomocephaloidea was short-lived and had a limited geographic distri ...
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Caudal Vertebra
The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic irregular bone whose complex structure is composed primarily of bone, and secondarily of hyaline cartilage. They show variation in the proportion contributed by these two tissue types; such variations correlate on one hand with the cerebral/caudal rank (i.e., location within the backbone), and on the other with phylogenetic differences among the vertebrate taxa. The basic configuration of a vertebra varies, but the bone is its ''body'', with the central part of the body constituting the ''centrum''. The upper (closer to) and lower (further from), respectively, the cranium and its central nervous system surfaces of the vertebra body support attachment to the intervertebral discs. The posterior part of a vertebra forms a vertebral arch (i ...
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Tiarajudens
''Tiarajudens'' (" Tiaraju tooth") is an extinct genus of saber-toothed herbivorous anomodonts which lived during the Middle Permian period ( Capitanian stage) in what is now Rio Grande do Sul, Brazil. It is known from the holotype UFRGS PV393P, a nearly complete skull. The type species ''T. eccentricus'' was named in 2011. Description ''Tiarajudens'' is a member of Anomodontia, a suborder of therapsids. Like other anomodonts, it was a quadrupedal herbivore about the size of a wild boar. The single fragmented holotype skull is short and robust at about in length. The most prominent features of ''Tiarajudens'' are its two large saber-like canine teeth. These teeth are unlike the tusks of dicynodonts, a later group of anomodonts. Twenty-one high- crowned teeth are present on either side of the upper jaw, including spoon-shaped incisors. Wide palatal teeth are also present. The top and bottom sets of teeth fit closely together, much like the teeth of mammals, allowing it ...
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