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Cave Insects
Cave-dwelling insects are among the most widespread and prominent troglofauna (cave-dwelling animals), including troglobites, troglophiles, and trogloxenes. As a category of ecological adaptations, such insects are significant in many senses, ecological, evolutionary, and physiological. Introduction A cave is an unusually well-defined ecological habitat in terms of its nature, time, and place. Accordingly, it is not surprising that a number of insects permanently inhabit caves, especially at the deepest levels, and are markedly specialised for niches in some of the extreme conditions. These are the true cavernicole species; troglobites rather than troglophiles or trogloxenes. These include; spiders and insects. Cavernicolous insect species rarely are adapted to move from cave to cave, so each species or community generally will be restricted wholly to certain caves or cave systems each, and commonly will have evolved in their respective home cave systems. Exceptions commonly are ...
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Troglofauna
Troglofauna are small cave-dwelling animals that have adapted to their dark surroundings. Troglofauna and stygofauna are the two types of subterranean fauna (based on life-history). Both are associated with subterranean environments – troglofauna are associated with caves and spaces above the water table and stygofauna with water. Troglofaunal species include spiders, insects, myriapods and others. Some troglofauna live permanently underground and cannot survive outside the cave environment. Troglofauna adaptations and characteristics include a heightened sense of hearing, touch and smell. Loss of under-used senses is apparent in the lack of pigmentation as well as eyesight in most troglofauna. Troglofauna insects may exhibit longer appendages and a lack of wings. Ecological categories Troglofauna are divided into three main categories based on their ecology: * Troglobionts (or troglobites): species, or populations of species, strictly bound to subterranean habitats. * Troglo ...
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Oligochaeta
Oligochaeta () is a subclass of soft-bodied animals in the phylum Annelida, which is made up of many types of aquatic and terrestrial worms, including all of the various earthworms. Specifically, oligochaetes comprise the terrestrial megadrile earthworms (some of which are semiaquatic or fully aquatic), and freshwater or semiterrestrial microdrile forms, including the tubificids, pot worms and ice worms ( Enchytraeidae), blackworms ( Lumbriculidae) and several interstitial marine worms. With around 10,000 known species, the Oligochaeta make up about half of the phylum Annelida. These worms usually have few setae (chaetae) or "bristles" on their outer body surfaces, and lack parapodia, unlike polychaeta. Diversity Oligochaetes are well-segmented worms and most have a spacious body cavity (coelom) used as a hydroskeleton. They range in length from less than up to in the 'giant' species such as the giant Gippsland earthworm (''Megascolides australis'') and the Mekong ...
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Acari
Mites are small arachnids (eight-legged arthropods) of two large orders, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari. However, most recent genetic analyses do not recover the two as each other's closest relative within Arachnida, rendering the group invalid as a clade. Most mites are tiny, less than in length, and have a simple, unsegmented body plan. The small size of most species makes them easily overlooked; some species live in water, many live in soil as decomposers, others live on plants, sometimes creating galls, while others are predators or parasites. This last type includes the commercially destructive '' Varroa'' parasite of honey bees, as well as scabies mites of humans. Most species are harmless to humans, but a few are associated with allergies or may transmit diseases. The scientific discipline devoted to the study of mites is called acarology. Evolution and taxonomy Mites are not a defined taxon, b ...
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Chilopoda
Centipedes (from Neo-Latin , "hundred", and Latin language, Latin , "foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', "lip", and Neo-Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, an arthropod group which includes millipedes and other multi-legged animals. Centipedes are elongated segmented (Metamerism (biology), metameric) animals with one pair of legs per body segment. All centipedes are venomous and can inflict painful centipede bite, stings, injecting their venom through Pincer (biology), pincer-like appendages known as forcipules or toxicognaths, which are actually modified legs instead of fangs. Despite the name, no species of centipede has exactly 100 legs; the number of pairs of legs is an odd number that ranges from 15 pairs to 191 pairs. Centipedes are predominantly generalist (biology), generalist carnivore, carnivorous, hunting for a variety of prey items that can be overpowered. They have ...
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Syncarida
Syncarida is a superorder of crustaceans, comprising the two extant order (biology), orders Anaspidacea and Bathynellacea, and the extinct order Palaeocaridacea. Taxonomy Fifty-nine living genera are known, in six families: ;Anaspidacea Calman, 1904 *Anaspidesidae Ahyong & Alonso-Zarazaga, 2017 *Koonungidae Sayce, 1908 *Psammaspididae Schminke, 1974 *Stygocarididae Noodt, 1963 ;Bathynellacea Chappuis, 1915 *Bathynellidae Grobben, 1904 *Parabathynellidae Noodt, 1965 References

* Syncarida, Arthropod superorders Taxa named by Alpheus Spring Packard {{Malacostraca-stub ...
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Isopoda
Isopoda is an order of crustaceans. Members of this group are called isopods and include both aquatic species and terrestrial species such as woodlice. All have rigid, segmented exoskeletons, two pairs of antennae, seven pairs of jointed limbs on the thorax, and five pairs of branching appendages on the abdomen that are used in respiration. Females brood their young in a pouch under their thorax called the marsupium. Isopods have various feeding methods: some eat dead or decaying plant and animal matter, others are grazers or filter feeders, a few are predators, and some are internal or external parasites, mostly of fish. Aquatic species mostly live on the seabed or the bottom of freshwater bodies of water, but some taxa can swim for short distance. Terrestrial forms move around by crawling and tend to be found in cool, moist places. Some species are able to roll themselves into a ball as a defense mechanism or to conserve moisture like species in the family Armadilli ...
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Decapoda
The Decapoda or decapods, from Ancient Greek δεκάς (''dekás''), meaning "ten", and πούς (''poús''), meaning "foot", is a large order of crustaceans within the class Malacostraca, and includes crabs, lobsters, crayfish, shrimp, and prawns. Most decapods are scavengers. The order is estimated to contain nearly 15,000 extant species in around 2,700 genera, with around 3,300 fossil species. Nearly half of these species are crabs, with the shrimp (about 3,000 species) and Anomura including hermit crabs, king crabs, porcelain crabs, squat lobsters (about 2500 species) making up the bulk of the remainder. The earliest fossils of the group date to the Devonian. Anatomy Decapods can have as many as 38 appendages, arranged in one pair per body segment. As the name Decapoda (from the Greek , ', "ten", and , '' -pod'', "foot") implies, ten of these appendages are considered legs. They are the pereiopods, found on the last five thoracic segments. In many decapods, one ...
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Copepoda
Copepods (; meaning 'oar-feet') are a group of small crustaceans found in nearly every freshwater and saltwater habitat. Some species are planktonic (living in the water column), some are benthic (living on the sediments), several species have parasitic phases, and some continental species may live in limnoterrestrial habitats and other wet terrestrial places, such as swamps, under leaf fall in wet forests, bogs, springs, ephemeral ponds, puddles, damp moss, or water-filled recesses of plants ( phytotelmata) such as bromeliads and pitcher plants. Many live underground in marine and freshwater caves, sinkholes, or stream beds. Copepods are sometimes used as biodiversity indicators. As with other crustaceans, copepods have a larval form. For copepods, the egg hatches into a nauplius form, with a head and a tail but no true thorax or abdomen. The larva molts several times until it resembles the adult and then, after more molts, achieves adult development. The nauplius form is so ...
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Cladocera
The Diplostraca or Cladocera, commonly known as water fleas, is a superorder (biology), superorder of small, mostly freshwater crustaceans, most of which feed on microscopic chunks of organic matter, though some forms are predatory. Over 1000 species have been recognised so far, with many more undescribed species, undescribed. The oldest fossils of diplostracans date to the Jurassic, though their modern morphology suggests that they originated substantially earlier, during the Paleozoic. Some have also adapted to a life in the ocean, the only members of Branchiopoda to do so, though several anostracans live in hypersaline lakes. Most are long, with a down-turned head with a single median compound eye, and a carapace covering the apparently unsegmented Crustacean#Structure, thorax and abdomen. Most species show cyclical parthenogenesis, where asexual reproduction is occasionally supplemented by sexual reproduction, which produces resting eggs that allow the species to survive hars ...
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Amphipoda
Amphipoda () is an order (biology), order of malacostracan crustaceans with no carapace and generally with laterally compressed bodies. Amphipods () range in size from and are mostly detritivores or scavengers. There are more than 10,700 amphipod species currently recognized. They are mostly marine animals, but are found in almost all aquatic environments. Some 2,250 species live in fresh water, and the order also includes the terrestrial Talitridae, sandhoppers such as ''Talitrus saltator'' and ''Arcitalitrus sylvaticus''. Etymology and names The name ''Amphipoda'' comes, via Neo-Latin ', from the Greek language, Greek root (linguistics), roots 'on both/all sides' and 'foot'. This contrasts with the related Isopoda, which have a single kind of thoracic leg. Particularly among Angling, anglers, amphipods are known as ''freshwater shrimp'', ''scuds'', or ''sideswimmers''. Description Anatomy The body of an amphipod is divided into 13 segments, which can be tagmosis, grouped ...
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Crustacea
Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthropods including decapods (shrimps, prawns, crabs, lobsters and crayfish), seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, opossum shrimps, amphipods and mantis shrimp. The crustacean group can be treated as a subphylum under the clade Mandibulata. It is now well accepted that the hexapods (insects and entognathans) emerged deep in the Crustacean group, with the completed pan-group referred to as Pancrustacea. The three classes Cephalocarida, Branchiopoda and Remipedia are more closely related to the hexapods than they are to any of the other crustaceans ( oligostracans and multicrustaceans). The 67,000 described species range in size from '' Stygotantulus stocki'' at , to the Japanese spider ...
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Fish
A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can be grouped into the more basal (phylogenetics), basal jawless fish and the more common jawed fish, the latter including all extant taxon, living cartilaginous fish, cartilaginous and bony fish, as well as the extinct placoderms and acanthodians. In a break to the long tradition of grouping all fish into a single Class (biology), class (Pisces), modern phylogenetics views fish as a paraphyletic group. Most fish are ectotherm, cold-blooded, their body temperature varying with the surrounding water, though some large nekton, active swimmers like white shark and tuna can hold a higher core temperature. Many fish can communication in aquatic animals#Acoustic, communicate acoustically with each other, such as during courtship displays. The stud ...
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