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Bungartiidae
''Bungartius'' is an extinct genus of arthrodire ("joint-necked") fish from the Late Devonian period of Ohio. It is the sole member of the family Bungartiidae, related to the giant ''Titanichthys''. The first specimen of ''Bungartius'' was recovered by Peter A. Bungart in Cuyahuga County, Ohio, and came from strata belonging to the Cleveland Shale, Cleveland Shale Member of the Ohio Shale. A second, more complete specimen was discovered in 1943 by David Dunkle, David H. Dunkle, who, four years later, named a new genus and species to accommodate them. ''Bungartius'' may have been Durophagy, durophagous, meaning that it would have eaten hard-shelled Invertebrate, invertebrates. Discovery In 1936, Peter Anthony Bungart, a staff member of the Cleveland Museum of Natural History, recovered a concretion containing the dorsal and ventral armour plates of an unknown fish from Cuyahoga County, Ohio, by Peter A. Bungart. The concretion originated from the Cleveland Shale, sometimes consi ...
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Late Devonian
The Devonian ( ) is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at million years ago ( Ma), to the beginning of the succeeding Carboniferous period at Ma. It is the fourth period of both the Paleozoic and the Phanerozoic. It is named after Devon, South West England, where rocks from this period were first studied. The first significant evolutionary radiation of life on land occurred during the Devonian, as free- sporing land plants (pteridophytes) began to spread across dry land, forming extensive coal forests which covered the continents. By the middle of the Devonian, several groups of vascular plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants ( pteridospermatophytes) appeared. This rapid evolution and colonization process, which had begun during the Silurian, is known as the Silurian-Devonian Terrestrial Rev ...
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Dunkleosteus
''Dunkleosteus'' is an extinct genus of large arthrodira, arthrodire ("jointed-neck") fish that existed during the Late Devonian period, about 382–358 million years ago. It was a pelagic fish inhabiting open waters, and one of the first vertebrate apex predators of any ecosystem. ''Dunkleosteus'' consists of ten species, some of which are among the largest placoderms ("plate-skinned") to have ever lived: ''D. terrelli'', ''D. belgicus'', ''D. denisoni'', ''D. marsaisi'', ''D. magnificus'', ''D. missouriensis'', ''D. newberryi'', ''D. amblyodoratus'', ''D. raveri'', and ''D. tuderensis.'' However, the validity of several of these species is unclear. The largest and best known species is ''D. terrelli''. Since body shape is not known, various methods of estimation put the living total length of the largest known specimen of ''D. terrelli'' between long and its weight around . Lengths of or more are poorly supported, with the most recent and extensive studies on the body shape a ...
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Bruntonichthys Multidens
''Bruntonichthys'' is an extinct monospecific genus of arthrodire placoderm from the Early Frasnian stage of the Late Devonian period. Fossils are found in the Gogo Formation of the Kimberley region of Australia. The skull is about 139 millimetres long, and had disproportionally large eye sockets. Researchers suggest it may have preyed on small mollusks.DENNIS, K. and MILES, R. S. (1980), New durophagous arthrodires from Gogo, Western Australia. Zoological Journal of the Linnean Society, 69: 43–85. Phylogeny ''Bruntonichthys'' is a basal member of the clade Aspinothoracidi, which belongs to the clade Pachyosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree or phylogeny is a graphical representation which shows the evolutionary history between a set of species or Taxon, taxa during a specific time.Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, M ... of ''Bruntonichth ...
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Trematosteus Fontanellus
''Trematosteus'' is an extinct monospecific genus of arthrodire placoderm from the Late Frasnian stage of the Late Devonian period. Fossils are found from Bad Wildungen, Germany. Phylogeny ''Trematosteus'' is a member of the superfamily Incisoscutoidea, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree or phylogeny is a graphical representation which shows the evolutionary history between a set of species or Taxon, taxa during a specific time.Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, M ... of ''Trematosteus'': References Arthrodires {{Placoderm-stub ...
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Tapinosteus Heintzi
''Tapinosteus'' is an extinct monospecific genus of arthrodire placoderm from the Late Frasnian stage of the Late Devonian period. Fossils are found from Bad Wildungen, Germany. Phylogeny ''Tapinosteus'' is a basal member of the clade Aspinothoracidi, which belongs to the clade Pachyosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree or phylogeny is a graphical representation which shows the evolutionary history between a set of species or Taxon, taxa during a specific time.Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, M ... of ''Tapinosteus'': References Arthrodires Arthrodire genera {{Placoderm-stub ...
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Rhachiosteus Pterygiatus
''Rhachiosteus'' is an extinct monospecific genus of arthrodire placoderm from the Middle to Late Devonian of Germany. It is known only from a single specimen, which may be a larval or juvenile form, as the skull of said specimen is only 19 millimetres long. Phylogeny ''Rhachiosteus'' is a basal member of the clade Pachyosteomorphi, the sister taxon to Coccosteomorphi, which together are the two main sub-clades of Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree or phylogeny is a graphical representation which shows the evolutionary history between a set of species or Taxon, taxa during a specific time.Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, M ... of ''Rhachiosteus'': References Arthrodires Placoderms of Europe Arthrodire genera {{Placoderm-stub ...
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Journal Of Paleontology
The ''Journal of Paleontology'' is a peer-reviewed scientific journal covering the field of paleontology. It is managed and published by Cambridge University Press on behalf of the Paleontological Society. Indexing The ''Journal of Paleontology'' is indexed in: *BIOSIS Previews *Science Citation Index *The Zoological Record *GeoRef __NOTOC__ The GeoRef database is a bibliographic database that indexes scientific literature in the geosciences, including geology. Coverage ranges from 1666 to the present for North American literature, and 1933 to the present for the rest of th ... References Paleontology journals Academic journals established in 1927 Academic journals published by learned and professional societies Cambridge University Press academic journals Bimonthly journals Paleontological Society {{paleo-journal-stub ...
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Phylogenetic
In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data and observed heritable traits of DNA sequences, protein amino acid sequences, and morphology. The results are a phylogenetic tree—a diagram depicting the hypothetical relationships among the organisms, reflecting their inferred evolutionary history. The tips of a phylogenetic tree represent the observed entities, which can be living taxa or fossils. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the taxa represented on the tree. An unrooted tree diagram (a network) makes no assumption about directionality of character state transformation, and does not show the origin or "root" of the taxa in question. In addition to their use for inferring phylogenetic pa ...
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Paraphyly
Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic grouping (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term received currency during the debates of the 1960s and 1970s accompanying the rise of cladistics, having been coined by zoologist Willi Hennig to apply to well-known taxa like Reptilia (reptiles), which is paraphyletic with respect to birds. Reptilia contains the last common ancestor of reptiles and all descendants of that ancest ...
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Pachyosteomorphi
Pachyosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci (of the suborder Brachythoraci), armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Pachyosteomorphi is the sister taxon to Coccosteomorphi, which together are the two main sub-clades of Eubrachythoraci. Pachyosteomorphi can be further sub-divided into Aspinothoracidi and Dunkleosteoidea, as shown in the cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ... below: References Arthrodires Devonian fish {{Placoderm-stub ...
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Diplognathus
''Diplognathus'' is a genus of arthrodire placoderm from the Late Famennian Cleveland Shale of Late Devonian Ohio, known only from incomplete fragments of jaws and skulls. What fragments are known suggest that the living animals were large-eyed piscivores with weak, but widely gaping jaws. ''D. mirabilis'' is thought to be fairly large, with infragnathals up to 45 centimeters in length. The second species, ''D. larfargei'', was much smaller, with inferognathals averaging about 4 centimeters in length. In 1967, Obruchev placed this genus within Hadrosteidae, on the basis of how the two genera have similar denticle ("teeth") patterns of the inferognathals, though Denison (1978) contested this placement, preferring to leave the taxon as Arthrodira ''incertae sedis or is a term used for a taxonomy (biology), taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system o ...
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