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Arthropleuridea
Arthropleuridea, from Ancient Greek ἄρθρον (''árthron''), meaning "joint", and πλευρά (''pleurá''), meaning "rib", is an extinct subclass of myriapod arthropods that flourished during the Carboniferous period, having first arisen during the Silurian, and perishing in the Early Permian. Members are characterized by possessing diplosegement (fused "double segments", as in modern-day millipedes) Paranota, paranotal tergal lobes separated from the body axis by a suture, and by Sclerosis (medicine), sclerotized plates buttressing the leg insertions. Despite their unique features, recent Phylogenetics, phylogenetic research suggests Arthropleuridea be included among millipedes in the class Diplopoda. The subclass contains three or two recognized Order (biology), orders, each with a single genus. Paleobiology Arthropleurids lived in the moist coal swamps that were common at the time and may have burrowed in the undergrowth. They were either herbivores or detritivores. Bes ...
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Millipedes
Millipedes (originating from the Latin , "thousand", and , "foot") are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a result of two single segments fused together. Most millipedes have very elongated cylindrical or flattened bodies with more than 20 segments, while pill millipedes are shorter and can roll into a tight ball. Although the name "millipede" derives from Latin for "thousand feet", no species was known to have 1,000 or more until the discovery in 2020 of '' Eumillipes persephone'', which can have over 1,300 legs. There are approximately 12,000 named species classified into 16 orders and around 140 families, making Diplopoda the largest class of myriapods, an arthropod group which also includes centipedes and other multi-legged creatures. Most millipedes are slow-moving detritivores, eating ...
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Myriapoda
Myriapods () are the members of subphylum Myriapoda, containing arthropods such as millipedes and centipedes. The group contains about 13,000 species, all of them terrestrial. Although molecular evidence and similar fossils suggests a diversification in the Cambrian Period, the oldest known fossil record of myriapods dates between the Late Silurian and Early Devonian, with '' Pneumodesmus'' preserving the earliest known evidence of air-breathing on land. Other early myriapod fossil species around the similar time period include '' Kampecaris obanensis'' and '' Archidesmus'' sp. The phylogenetic classification of myriapods is still debated. The scientific study of myriapods is myriapodology, and those who study myriapods are myriapodologists. Anatomy Myriapods have a single pair of antennae and, in most cases, simple eyes. Exceptions are the two classes of symphylans and pauropods, the millipede order Polydesmida and the centipede order Geophilomorpha, which are all ey ...
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Eoarthropleura
''Eoarthropleura'' was a genus of millipede-like creatures which lived between the Late Silurian and Late Devonian periods. It reached in length. Fossils, mainly of cuticle fragments, have been found in Europe (Rhineland-Palatinate, Germany and Shropshire, England) and North America (New York, USA and New Brunswick, Canada). It is the earliest known member of the Arthropleuridea Arthropleuridea, from Ancient Greek ἄρθρον (''árthron''), meaning "joint", and πλευρά (''pleurá''), meaning "rib", is an extinct subclass of myriapod arthropods that flourished during the Carboniferous period, having first arisen d ..., and the oldest known terrestrial animal of North America. References Silurian myriapods Devonian myriapods Late Devonian animals Pridoli first appearances Frasnian extinctions Paleozoic arthropods of North America Paleozoic arthropods of Europe Prehistoric myriapod genera Fossil taxa described in 1976 Millipedes {{paleo-myriapod- ...
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Arthropleura
''Arthropleura'', from Ancient Greek ἄρθρον (''árthron''), meaning "joint", and πλευρά (''pleurá''), meaning "rib", is an extinct genus of massive myriapoda, myriapod that lived in what is now Europe and North America around 344 to 292 Myr, million years ago, from the Viséan stage of the lower Carboniferous period to the Sakmarian stage of the lower Permian period. It is a millipede, and was capable of reaching at least in length, possibly up to over , making it the largest known land arthropod of all time. ''Arthropleura'' is known from body fossils as well as trace fossils, particularly giant trackways up to wide, and potentially also large burrows. It lived in open, sparsely wooded environments near water, and was possibly amphibious. Classification First discovered in 1849 during construction of a railway near Friedrichsthal in Germany, ''Arthropleura'' has consistently attracted much artistic and scientific attention, yet has historically been known fr ...
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Detritivore
Detritivores (also known as detrivores, detritophages, detritus feeders or detritus eaters) are heterotrophs that obtain nutrients by consuming detritus (decomposing plant and animal parts as well as feces). There are many kinds of invertebrates, vertebrates, and plants that eat detritus or carry out coprophagy. By doing so, all these detritivores contribute to decomposition and the nutrient cycles. Detritivores should be distinguished from other decomposers, such as many species of bacteria, fungi and protists, which are unable to ingest discrete lumps of matter. Instead, these other decomposers live by absorbing and metabolizing on a molecular scale (saprotrophic nutrition). The terms ''detritivore'' and ''decomposer'' are often used interchangeably, but they describe different organisms. Detritivores are usually arthropods and help in the process of remineralization. Detritivores perform the first stage of remineralization, by fragmenting the dead plant matter, allowing decomp ...
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Exoskeleton
An exoskeleton () . is a skeleton that is on the exterior of an animal in the form of hardened integument, which both supports the body's shape and protects the internal organs, in contrast to an internal endoskeleton (e.g. human skeleton, that of a human) which is enclosed underneath other soft tissues. Some large, hard and non-flexible protective exoskeletons are known as mollusc shell, shell or armour (anatomy), armour. Examples of exoskeletons in animals include the arthropod exoskeleton, cuticle skeletons shared by arthropods (insects, chelicerates, myriapods and crustaceans) and tardigrades, as well as the corallite, skeletal cups formed by hardened secretion of stony corals, the test (biology), test/tunic of sea squirts and sea urchins, and the prominent mollusc shell shared by snails, bivalvia, clams, tusk shells, chitons and nautilus. Some vertebrate animals, such as the turtle, have both an endoskeleton and a turtle shell, protective exoskeleton. Role Exoskeletons c ...
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Spiracle (arthropods)
A spiracle or stigma is the opening in the exoskeletons of insects, myriapods, velvet worms and many arachnids to allow air to enter the trachea. Insect respiratory system differs from vertebrates'. The circulatory system plays a relatively minor role in circulating oxygen and removing carbon dioxide; instead, trachea and air sacs in the insect body allow direct gas exchange, and these tracheal tubes eventually connect to the external environment via spiracles. In most species, the spiracles are controlled by motor neurons in the central nervous system. It can be opened and closed in an efficient manner to admit air while minimizing associated physiological costs, such as water loss during respiration. Many sensory stimuli can affect the control of spiracles in insects, e.g. chemosensory (carbon dioxide, oxygen, etc.) or mechanosensory (sound, touch, etc.). It has been shown that during metabolically intensive behaviors, such as flight, insects can dynamically modulate the spir ...
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Lung
The lungs are the primary Organ (biology), organs of the respiratory system in many animals, including humans. In mammals and most other tetrapods, two lungs are located near the Vertebral column, backbone on either side of the heart. Their function in the respiratory system is to extract oxygen from the atmosphere and transfer it into the bloodstream, and to release carbon dioxide from the bloodstream into the atmosphere, in a process of gas exchange. Respiration is driven by different muscular systems in different species. Mammals, reptiles and birds use their musculoskeletal systems to support and foster breathing. In early tetrapods, air was driven into the lungs by the pharyngeal muscles via buccal pumping, a mechanism still seen in amphibians. In humans, the primary muscle that drives breathing is the Thoracic diaphragm, diaphragm. The lungs also provide airflow that makes Animal communication#Auditory, vocalisation including speech possible. Humans have two lungs, a ri ...
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Silurian
The Silurian ( ) is a geologic period and system spanning 23.5 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the third and shortest period of the Paleozoic Era, and the third of twelve periods of the Phanerozoic Eon. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods ( myriapods, arachnids and hexapods) ...
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Gill
A gill () is a respiration organ, respiratory organ that many aquatic ecosystem, aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they are kept moist. The microscopic structure of a gill presents a large surface area to the external environment. Branchia (: branchiae) is the zoologists' name for gills (from Ancient Greek ). With the exception of some aquatic insects, the filaments and lamella (surface anatomy), lamellae (folds) contain blood or Coelom#Coelomic fluid, coelomic fluid, from which gases are exchanged through the thin walls. The blood carries oxygen to other parts of the body. Carbon dioxide passes from the blood through the thin gill tissue into the water. Gills or gill-like organs, located in different parts of the body, are found in various groups of aquatic animals, including Mollusc, molluscs, crustaceans, insects, fish, a ...
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Aquatic Respiration
Aquatic respiration is the Biological process, process whereby an aquatic animal, aquatic organism exchanges Respiration (physiology), respiratory gases with water, obtaining oxygen from oxygen dissolved in water and excretion, excreting carbon dioxide and some other metabolic waste products into the water. Unicellular and simple small organisms In very small animals, plants and bacteria, simple diffusion of gaseous metabolites is sufficient for respiratory function and no special adaptations are found to aid respiration. Passive diffusion or active transport are also sufficient mechanisms for many larger aquatic animals such as many worms, jellyfish, sponges, bryozoans and similar organisms. In such cases, no specific respiratory organs or organelles are found. Higher plants Although higher plants typically use carbon dioxide and excrete oxygen during photosynthesis, they also respire and, particularly during darkness, many plants excrete carbon dioxide and require oxygen to ma ...
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