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Arthrodire
Arthrodira (Greek for "jointed neck") is an Order (biology), order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are covered by a Sclerotic ring, bony ring, which supports the eye, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus ''Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, ''Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In ...
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Dunkleosteus Terrelli
''Dunkleosteus'' is an extinct genus of large arthrodira, arthrodire ("jointed-neck") fish that existed during the Late Devonian period, about 382–358 million years ago. It was a pelagic fish inhabiting open waters, and one of the first vertebrate apex predators of any ecosystem. ''Dunkleosteus'' consists of ten species, some of which are among the largest placoderms ("plate-skinned") to have ever lived: ''D. terrelli'', ''D. belgicus'', ''D. denisoni'', ''D. marsaisi'', ''D. magnificus'', ''D. missouriensis'', ''D. newberryi'', ''D. amblyodoratus'', ''D. raveri'', and ''D. tuderensis.'' However, the validity of several of these species is unclear. The largest and best known species is ''D. terrelli''. Since body shape is not known, various methods of estimation put the living total length of the largest known specimen of ''D. terrelli'' between long and its weight around . Lengths of or more are poorly supported, with the most recent and extensive studies on the body shape a ...
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Placodermi
Placoderms (from Ancient Greek πλάξ 'plax'', ''plakos''' plate' and δέρμα 'derma'''skin') are vertebrate animals of the class Placodermi, an extinct group of prehistoric fish known from Paleozoic fossils during the Silurian and the Devonian periods. While their endoskeletons are mainly cartilaginous, their head and thorax were covered by articulated armoured plates (hence the name), and the rest of the body was scaled or naked depending on the species. Placoderms were among the first jawed fish (their jaws likely evolved from the first pair of gill arches), as well as the first vertebrates to have true teeth. They were also the first fish clade to develop pelvic fins, the second set of paired fins and the homologous precursor to hindlimbs in tetrapods. 380-million-year-old fossils of three other genera, '' Incisoscutum'', '' Materpiscis'' and '' Austroptyctodus'', represent the oldest known examples of live birth. Placoderms are thought to be paraphyletic ...
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Arthrodira
Arthrodira (Greek for "jointed neck") is an order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are covered by a bony ring, which supports the eye, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus '' Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, ''Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In contrast, the long-nosed '' Ro ...
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Incisoscutum
''Incisoscutum'' is an extinct genus of arthrodire placoderm from the Early Frasnian Gogo Reef, from Late Devonian Australia. The genus contains two species ''I. ritchiei'', named after Alex Ritchie, a ichthyology, palaeoichthyologist and senior fellow of the Australian Museum, and ''I. sarahae'', named after Sarah Long, daughter of its discoverer and describer, John A. Long. The genus is important in the study of early vertebrates as well-preserved fossilized embryos have been found in female specimens and ossified pelvic claspers found in males. This shows that viviparity and internal fertilization was common amongst these primitive jawed vertebrates, which are outside the crown group Gnathostomata. In a study of fossil remains, comparison of the ontogeny of fourteen dermal plates from ''Compagopiscis, Compagopiscis croucheri'' and the more derived species ''Incisoscutum ritchiei'' suggested that lengthwise growth occurs earlier in the ontogeny than growth in width, and that di ...
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Phlyctaenioidei
Arthrodira (Greek for "jointed neck") is an order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are covered by a bony ring, which supports the eye, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus '' Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, ''Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In contrast, the long-nosed '' Ro ...
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Groenlandaspididae
Arthrodira (Greek for "jointed neck") is an Order (biology), order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are covered by a Sclerotic ring, bony ring, which supports the eye, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus ''Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, ''Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In ...
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Arctolepididae
Arthrodira (Greek for "jointed neck") is an order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are covered by a bony ring, which supports the eye, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus '' Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, ''Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In contrast, the long-nosed '' Ro ...
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Rolfosteus
''Rolfosteus'' is an extinct monospecific genus of arthrodire placoderm from the Early Frasnian stage of the Late Devonian period, found at the Gogo Formation of Western Australia. Description ''Rolfosteus'' had a skull reaching , with tough plating on the front of its body. Like other arthrodires, such as ''Coccosteus'' and the giant ''Dunkleosteus'', it had sharp, bony plates on its mouth which formed a turtle-like beak for cutting prey to pieces. ''Rolfosteuss most unusual feature was its highly elongated rostrum (snout), which may have been used to enhance its sense of smell as well as increase its hydrodynamic streamlining, making it well adapted to be a pelagic pursuit predator. Its fast jaw movement and jaw morphology allowed it to effectively capture prey and swallow them whole. Phylogeny ''Rolfosteus'' is a member of the family Camuropiscidae under the superfamily Incisoscutoidea, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachy ...
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Actinolepidae
Actinolepidae is an extinct family of placoderm fishes which lived during the Early Devonian period. They are considered to be among the most primitive of the arthrodires, and are widely accepted to be phylogenetically basal to the group. Description The bodies of Actinolepids are wide and flat, suggesting that most members of this family were benthic fish. Their jaws were comparatively underdeveloped in comparison to the more robust-jawed arthrodires that would come after them, such as ''Dunkleosteus ''Dunkleosteus'' is an extinct genus of large arthrodira, arthrodire ("jointed-neck") fish that existed during the Late Devonian period, about 382–358 million years ago. It was a pelagic fish inhabiting open waters, and one of the first verteb ...'' and '' Coccosteus'', indicating that it likely subsisted primarily on smaller, softer-bodied animals such as mollusks, or worms instead of larger, tougher prey animals. References External links * Devonian placoderms ...
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Arctolepis
''Arctolepis'' is an extinct genus of placoderm arthrodire fish which lived during the Early Devonian period (408 – 307 million years ago). Fossils of ''Arctolepis'' have been found in what is now Norway and Michigan. Classification Within Arthrodira, ''Arctolepis'' is considered a member of "Phylctaenii", a paraphyletic grouping within the clade Phlyctaenioidei, basal to the suborder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized ... Brachythoraci. ''Arctolepis'' is most closely related to '' Heintzosteus'' and '' Dicksonosteus''. ''Arctolepis'' contains the following six species: * ''Arctolepis brevis'' Heintz, 1929 * ''Arctolepis decipiens'' Woodward, 1891 * ''Arctolepis lata'' Heintz, 1929 * ''Arctolepis lewini'' Heintz, 1929 * ''Arctolepis longicornis'' Heintz, 1929 * ''A ...
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Devonian
The Devonian ( ) is a period (geology), geologic period and system (stratigraphy), system of the Paleozoic era (geology), era during the Phanerozoic eon (geology), eon, spanning 60.3 million years from the end of the preceding Silurian period at million years ago (Megaannum, Ma), to the beginning of the succeeding Carboniferous period at Ma. It is the fourth period of both the Paleozoic and the Phanerozoic. It is named after Devon, South West England, where rocks from this period were first studied. The first significant evolutionary radiation of history of life#Colonization of land, life on land occurred during the Devonian, as free-spore, sporing land plants (pteridophytes) began to spread across dry land, forming extensive coal forests which covered the continents. By the middle of the Devonian, several groups of vascular plants had evolved leaf, leaves and true roots, and by the end of the period the first seed-bearing plants (Pteridospermatophyta, pteridospermatophyt ...
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