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Archipolypoda Cross-section
Archipolypoda is an extinct group of millipedes known from fossils in Europe and North America and containing the earliest known land animals. The Archipolypoda was erected by Scudder (1882) but redefined in 2005 with the description of several new species from Scotland. Distinguishing characteristics include relatively large eyes with densely packed ocelli (sometimes interpreted as compound eyes), and modified leg pairs on the 8th body ring. Some species had prominent spines while others had a flattened appearance. Classification The Archipolypoda as currently recognized consists of four orders, many with monotypic families and genera, as well as five species of uncertain placement (''incertae sedis''). Archidesmida Wilson & Anderson, 2004 * Archidesmidae Scudder, 1885 **'' Archidesmus macnicoli'' Peach, 1882. Lower Devonian, Scotland * Zanclodesmidae Wilson, Daeschler & Desbiens, 2005 **'' Zanclodesmus willetti'' Wilson, Daeschler & Desbiens, 2005. Upper Devonian, Queb ...
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Samuel Hubbard Scudder
Samuel Hubbard Scudder (April 13, 1837 – May 17, 1911) was an American entomologist and paleontologist. He was a leading figure in entomology during his lifetime and the founder of insect paleontology in America. In addition to fossil insects, he was an authority on butterflies ( Lepidoptera) and grasshoppers (Orthoptera). Biography Scudder was born on April 13, 1837, in Boston, Massachusetts, the son of Charles Scudder and Sarah Lathrop (Coit) Scudder. His father was a successful merchant, and both parents had Puritan roots dating back to the founding of the Massachusetts Bay Colony in the 1620s. He was raised in a strict Calvinist Congregational household.Leach (2013) One of his younger brothers, Horace Scudder, became a noted author and editor of the ''Atlantic Monthly'',Cockerell (1911) while his niece Vida Dutton Scudder was a writer and social activist. Scudder attended Boston Latin School, and then enrolled in Williams College in 1853 at the age of 16. He studied with ...
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Lower Devonian
The Early Devonian is the first of three epochs comprising the Devonian period, corresponding to the Lower Devonian series. It lasted from and began with the Lochkovian Stage , which was followed by the Pragian from and then by the Emsian, which lasted until the Middle Devonian began, . During this time, the first ammonoids appeared, descending from bactritoid nautiloids. Ammonoids during this time period were simple and differed little from their nautiloid counterparts. These ammonoids belong to the order Agoniatitida, which in later epochs evolved to new ammonoid orders, for example Goniatitida and Clymeniida. This class of cephalopod molluscs would dominate the marine fauna until the beginning of the Mesozoic The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Creta ... Era. Refer ...
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Euphoberia
''Euphoberia'' is an extinct genus of millipede from the Pennsylvanian epoch of the Late Carboniferous, measuring up to in length, that is small in Euphoberiidae, with length of largest species about . Fossils have been found in Europe and North America. There has been uncertainty about the appropriate classification of ''Euphoberia'' since its description in 1868: it has been referred to as a centipede, millipede, or a separate, independent group within the myriapods. It is currently placed in the Archipolypoda Archipolypoda is an extinct group of millipedes known from fossils in Europe and North America and containing the earliest known land animals. The Archipolypoda was erected by Scudder (1882) but redefined in 2005 with the description of seve ..., an extinct group of millipedes. Several species described in the late 19th century have since been assigned to the related genera '' Myriacantherpestes'' and '' Acantherpestes''. References Carboniferous myri ...
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Upper Carboniferous
Upper may refer to: * Shoe upper or ''vamp'', the part of a shoe on the top of the foot * Stimulant, drugs which induce temporary improvements in either mental or physical function or both * ''Upper'', the original film title for the 2013 found footage film ''The Upper Footage ''The Upper Footage'' (also known as ''Upper'') is a 2013 found footage film written and directed by Justin Cole. First released on January 31, 2013 to a limited run of midnight theatrical screenings at Landmark’s Sunshine Cinema in New York Cit ...'' See also

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Euphoberiidae
Euphoberiidae is an extinct family of archipolypodan millipedes known from the Upper Pennsylvanian of Europe and North America. The family includes relatively large millipedes measuring up to long bearing distinctive spines and were the dominant millipedes of the Pennsylvanian.Shear, William A. (1997The fossil record and evolution of the Myriapoda In: Fortey, R. A., Thomas, R. H. (Eds.), ''Arthropod Relationships''. Systematics Association Special Volume 55. Chapman & Hall, London, pp. 211–219 Description Euphoberiids are characterized by sets of outward-pointing spines on each diplosegment (body ring), one pair extending laterally (to the sides), the other pair higher on the body. At the base of each lateral spine is an ozopore An ozopore is the opening of a defensive gland present in some arthropods, notably in millipedes of the order Polydesmida
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Silurian
The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago (Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods ( myriapods, arachnids and hexapods) became fully terrestrialized. A significant evolutionary milestone d ...
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Cowiedesmus
''Cowiedesmus'' is an extinct millipede genus known from the middle Silurian of Scotland, and is one of the earliest known land animals. ''Cowiedesmus'' was about 4 cm (1.6 in) long and characterized by a greatly enlarged pair of legs on the 8th segment which may have been used in clasping females or functioned as gonopods (modified legs used to insert sperm during mating). ''Coweiedesmus'' is distinct enough from other living and fossil millipedes to be placed in its own order, Cowiedesmida. The only known species, ''C. eroticopodus'', was described in 2004. The name ''Cowiedesmus'' references the Cowie Formation, a geological formation at Cowie Harbor, near the village of Stonehaven Stonehaven ( , ) is a town in Scotland. It lies on Scotland's northeast coast and had a population of 11,602 at the 2011 Census. After the demise of the town of Kincardine, which was gradually abandoned after the destruction of its royal cast ..., Scotland, where the fossil was fo ...
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Pennsylvania
Pennsylvania (; (Pennsylvania Dutch: )), officially the Commonwealth of Pennsylvania, is a state spanning the Mid-Atlantic, Northeastern, Appalachian, and Great Lakes regions of the United States. It borders Delaware to its southeast, Maryland to its south, West Virginia to its southwest, Ohio to its west, Lake Erie and the Canadian province of Ontario to its northwest, New York to its north, and the Delaware River and New Jersey to its east. Pennsylvania is the List of U.S. states and territories by population, fifth-most populous state in the nation with over 13 million residents 2020 United States census, as of 2020. It is the List of U.S. states and territories by area, 33rd-largest state by area and ranks List of states and territories of the United States by population density, ninth among all states in population density. The southeastern Delaware Valley metropolitan area comprises and surrounds Philadelphia, the state's List of cities in Pennsylvania, largest ...
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Quebec, Canada
Quebec ( ; )According to the Canadian government, ''Québec'' (with the acute accent) is the official name in Canadian French and ''Quebec'' (without the accent) is the province's official name in Canadian English is one of the thirteen provinces and territories of Canada. It is the largest province by area and the second-largest by population. Much of the population lives in urban areas along the St. Lawrence River, between the most populous city, Montreal, and the provincial capital, Quebec City. Quebec is the home of the Québécois nation. Located in Central Canada, the province shares land borders with Ontario to the west, Newfoundland and Labrador to the northeast, New Brunswick to the southeast, and a coastal border with Nunavut; in the south it borders Maine, New Hampshire, Vermont, and New York in the United States. Between 1534 and 1763, Quebec was called ''Canada'' and was the most developed colony in New France. Following the Seven Years' War, Quebec bec ...
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