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Apankura
''Apankura'' is an extinct genus of Cambrian euthycarcinoids from the Santa Rosita Formation of Argentina. The genus contains a single species, ''Apankura machu''. It was at one point the only Cambrian euthycarcinoid. However, '' Mosineia'' and '' Mictomerus'' are likely older. Description ''Apankura'' is roughly long, and has large mandibles, a trait previously unknown from euthycarcinoids. The antennae are uniramous like other euthycarcinoids, with a possible buccal complex present behind the mandibles. No post-mandibular oral appendages are present. The pre-abdomen is composed of five tergites, with 11 uniramous limb pairs, with the tergites showing "segment decoupling" where there are more limbs than tergites. The pre-abdomen also has 11 sternites, each roughly 3 times wider than long. The pre-abdominal limbs match other euthycarcinoids, although they taper distally and lack setae. The apodemes are preserved as several dark rods. The post-abdomen is incomplete but co ...
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Euthycarcinoid
Euthycarcinoidea are an enigmatic group of extinct, possibly amphibious arthropods that ranged from Cambrian to Triassic times. Fossils are known from Europe, North America, Argentina, Australia, and Antarctica. Description The euthycarcinoid body was divided into a cephalon (head), preabdomen, and postabdomen. The cephalon consisted of two segments and included mandibles, antennae and presumed eyes. The preabdomen consisted of five to fourteen tergites, each having up to three somites. Each somite had in turn a pair of uniramous, segmented legs. The postabdomen was limbless and consisted of up to six segments and a terminal tail spine. Affinities Due to its particular combination of characteristics, the position of the Euthycarcinoidea within the Arthropoda has been ambiguous; previous authors have allied euthycarcinoids with crustaceans (interpreted as copepods, branchiopods, or an independent group), with trilobites, or the merostomatans ( horseshoe crabs and sea scorpi ...
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Cambrian
The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding Ediacaran period 538.8 Ma (million years ago) to the beginning of the Ordovician Period 486.85 Ma. Most of the continents lay in the southern hemisphere surrounded by the vast Panthalassa Ocean. The assembly of Gondwana during the Ediacaran and early Cambrian led to the development of new convergent plate boundaries and continental-margin arc magmatism along its margins that helped drive up global temperatures. Laurentia lay across the equator, separated from Gondwana by the opening Iapetus Ocean. The Cambrian marked a profound change in life on Earth; prior to the Period, the majority of living organisms were small, unicellular and poorly preserved. Complex, multicellular organisms gradually became more common during the Ediacaran, but it was not until the Cambrian that fossil diversity seems to rapidly ...
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Paleozoic Arthropods Of South America
The Paleozoic ( , , ; or Palaeozoic) Era is the first of three geological eras of the Phanerozoic Eon. Beginning 538.8 million years ago (Ma), it succeeds the Neoproterozoic (the last era of the Proterozoic Eon) and ends 251.9 Ma at the start of the Mesozoic Era. The Paleozoic is subdivided into six geologic periods (from oldest to youngest), Cambrian, Ordovician, Silurian, Devonian, Carboniferous and Permian. Some geological timescales divide the Paleozoic informally into early and late sub-eras: the Early Paleozoic consisting of the Cambrian, Ordovician and Silurian; the Late Paleozoic consisting of the Devonian, Carboniferous and Permian. The name ''Paleozoic'' was first used by Adam Sedgwick (1785–1873) in 1838 to describe the Cambrian and Ordovician periods. It was redefined by John Phillips (1800–1874) in 1840 to cover the Cambrian to Permian periods. It is derived from the Greek ''palaiós'' (παλαιός, "old") and ''zōḗ'' (ζωή, "life") meaning "ancient lif ...
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Fossil Taxa Described In 2004
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Though the fossil record is incomplete, numerous studies have demonstrated that there is enough information available to give a good understanding of the pattern of diversification of life on Earth. In addition, the record can predict and fill gaps such as the discovery of ''Tiktaalik'' in the arctic of Canada. Paleontology includes the study of fossils: their age, method of formation, and evolutionary significance. Specimens are sometimes considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years to 4.1 billion years old. Early edition, published online before print. The ob ...
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Gonopore
A gonopore, sometimes called a gonadopore, is a genital pore in many invertebrates. Hexapods, including insects, have a single common gonopore, except mayflies, which have a pair of gonopores. More specifically, in the unmodified female, it is the opening of the common oviduct, and in the male, it is the opening of the ejaculatory duct. The position of the gonopore varies considerably between groups, but is generally constant within groups, allowing its position to be used as a "segmental marker". In Malacostraca, it is on the sixth thoracic segment; in Symphyla, it is on the fourth trunk segment; in arachnids, it is on the second segment of the opisthosoma. In insects and centipedes, the gonopores are close to the animal's tail, while in millipede Millipedes (originating from the Latin , "thousand", and , "foot") are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Dipl ...
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Apodeme
An exoskeleton () . is a skeleton that is on the exterior of an animal in the form of hardened integument, which both supports the body's shape and protects the internal organs, in contrast to an internal endoskeleton (e.g. that of a human) which is enclosed underneath other soft tissues. Some large, hard and non-flexible protective exoskeletons are known as shell or armour. Examples of exoskeletons in animals include the cuticle skeletons shared by arthropods (insects, chelicerates, myriapods and crustaceans) and tardigrades, as well as the skeletal cups formed by hardened secretion of stony corals, the test/tunic of sea squirts and sea urchins, and the prominent mollusc shell shared by snails, clams, tusk shells, chitons and nautilus. Some vertebrate animals, such as the turtle, have both an endoskeleton and a protective exoskeleton. Role Exoskeletons contain rigid and resistant components that fulfil a set of functional roles in addition to structural support in ma ...
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Sternite
The sternum (: sterna) is the ventral portion of a segment of an arthropod thorax or abdomen. In insects, the sterna are usually single, large sclerites, and external. However, they can sometimes be divided in two or more, in which case the subunits are called sternites, and may also be modified on the terminal abdominal segments so as to form part of the functional genitalia, in which case they are frequently reduced in size and development, and may become internalized and/or membranous. For a detailed explanation of the terminology, see Kinorhynchs have tergal and sternal plates too, though seemingly not homologous with those of arthropods.Sørensen, M. V. et al. Phylogeny of Kinorhyncha based on morphology and two molecular loci. PLoS One 10, 1–33 (2015). Ventrites are externally visible sternites. Usually the first sternite is covered up, so that ventrite numbers do not correspond to sternite numbers. The term is also used in other arthropod groups such as crustaceans, ...
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Antenna (biology)
An antenna (plural: antennae) is one of a pair of appendages used for Sensory system, sensing in arthropods. Antennae are sometimes referred to as ''feelers''. Antennae are connected to the first one or two Segmentation (biology), segments of the arthropod head. They vary widely in form but are always made of one or more jointed segments. While they are typically sensory organs, the exact nature of what they sense and how they sense it is not the same in all groups. Functions may variously include sensing tactition, touch, air motion, heat, vibration (sound), and especially insect olfaction, smell or gustation, taste. Antennae are sometimes modified for other purposes, such as mating, brooding, swimming, and even anchoring the arthropod to a substrate (biology), substrate. Larval arthropods have antennae that differ from those of the adult. Many crustaceans, for example, have free-swimming larvae that use their antennae for swimming. Antennae can also locate other group members i ...
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Tergites
A ''tergum'' (Latin for "the back"; : ''terga'', associated adjective tergal) is the dorsal ('upper') portion of an arthropod segment other than the head. The anterior edge is called the 'base' and posterior edge is called the 'apex' or 'margin'. A given tergum may be divided into hardened plates or sclerites commonly referred to as tergites. In a thoracic segment, for example, the tergum may be divided into an anterior notum and a posterior scutellum. Lateral extensions of a tergite are known as paranota (Greek for "alongside the back") or ''carinae'' (Latin for "keel"), exemplified by the flat-backed millipedes of the order Polydesmida. Kinorhynchs have tergal and sternal plates too, though seemingly not homologous with those of arthropods. Tergo-tergal is a stridulatory mechanism in which fine spines of the abdominal tergites are rubbed together to produce sound. This process is known as abdominal telescoping. Examples File:Andrena spiraeana abdomen.jpg , Abdominal tergum ...
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Mandible (arthropod Mouthpart)
250px, The mandibles of a bull ant The mandible (from or mandĭbŭ-lum, a jaw) of an arthropod is a pair of mouthparts used either for biting or cutting and holding food. Mandibles are often simply called jaws. Mandibles are present in the extant subphyla Myriapoda (millipedes and others), Crustacea and Hexapoda (insects etc.). These groups make up the clade Mandibulata, which is currently believed to be the sister group to the rest of arthropods, the clade Arachnomorpha (Chelicerata and Trilobita). Unlike the chelicerae of arachnids, mandibles can often be used to chew food. Mandibulates also differ by having antennae, and also by having three distinct body regions: head, thorax and abdomen. (The cephalothorax (or prosoma) of chelicerates is a fusion of head and thorax.) Insects Insect mandibles are as diverse in form as their food. For instance, grasshoppers and many other plant-eating insects have sharp-edged mandibles that move side to side. Most butterflies ...
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Mictomerus
''Mictomerus'' is an extinct genus of Cambrian euthycarcinoids from the Potsdam Group of Quebec, Canada. The genus contains a single species, ''Mictomerus melochevillensis'', known from 29 specimens, all but one preserved ventrally. Description ''Mictomerus'' cephalic area is poorly known, with an elongate oval structure at the anterior portion with a narrow medial invagination, and two circular structures which may be eyes. The thorax has five to six overlapping tergites A ''tergum'' (Latin for "the back"; : ''terga'', associated adjective tergal) is the dorsal ('upper') portion of an arthropod segment other than the head. The anterior edge is called the 'base' and posterior edge is called the 'apex' or 'margin'. ..., with varying posterolateral margins. Due to the ventral preservation of the specimens, their dorsal surfaces are unknown. The thorax's segmentation is also unknown, however it can be inferred from the limbs. The abdomen is composed of five limbless segments ...
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