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Alienopteridae
Alienopteridae is an extinct family of dictyopterans, known from the Mid-Cretaceous to Eocene. They are noted for their unusual combination of features not found in other dictyopterans. Taxonomy It was originally assigned to its own order Alienoptera by Bai et al., 2016. It was reassigned to the dictyopteran superfamily Umenocoleoidea as sister family to the beetle-like Umenocoleidae by Vršanský ''et al.'' (2018), and a more recent analysis similarly places Alienopteridae and Umenocoleidae as sister taxa within Dictyoptera, but placing both lineages outside of Blattodea. A 2021 study revived the order Alienoptera for the clade containing Alienopteridae and Umenocoleidae, with a cladistic analysis placing Alienoptera as the sister clade to Mantodea (praying mantises). Distribution The majority of the alienopterid genera are known from the mid Cretaceous (latest Albian-earliest Cenomanian ~ 100 million years ago) Burmese amber found in Myanmar; though an additional two ...
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Umenocoleoidea
Dictyoptera (from Greek δίκτυον ''diktyon'' "net" and πτερόν ''pteron'' "wing") is an insect superorder that includes two extant orders of polyneopterous insects: the order Blattodea (termites and cockroaches together) and the order Mantodea (mantises). While all modern Dictyoptera have short ovipositors, the oldest fossils of Dictyoptera have long ovipositors, much like members of the Orthoptera. Classification and phylogeny The use of the term Dictyoptera has changed over the years, and while largely out of use for much of the last century, it is becoming more widely used. It has usually been considered a superorder, with Isoptera, Blattodea and Mantodea being its three orders. In some classifications, however, Dictyoptera is shifted to order status and in others the order Isoptera has been subsumed under Blattodea while retaining Dictyoptera as a superorder. Regardless, in all classifications the constituent groups are the same, just treated at different rank. T ...
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Blattodea
Blattodea is an order of insects that contains cockroaches and termites. Formerly, termites were considered a separate order, Isoptera, but genetic and molecular evidence suggests they evolved from within the cockroach lineage, cladistically making them cockroaches as well. The Blattodea and the mantis (order Mantodea) are now all considered part of the superorder Dictyoptera. Blattodea includes approximately 4,400 species of cockroach in almost 500 genera, and about 3,000 species of termite in around 300 genera. Termites are pale-coloured, soft-bodied eusocial insects that live in colonies, whereas cockroaches are darker-coloured (often brown), sclerotized, segmented insects. Within the colony, termites have a caste system, with a pair of mature reproductives, the king and the queen, and numerous sterile workers and soldiers. Cockroaches are not colonial but do have a tendency to aggregate and may be considered pre-social, as all adults are capable of breeding. Other simil ...
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Umenocoleidae
Umenocoleidae is an extinct family of dictyopteran insects known from the Cretaceous. They are considered to be closely related to the Alienopteridae. They were originally considered to be beetles due to their beetle-like morphology, with sclerotised elytra-like forewings. This was probably an adaptation for living under bark and in other tight spaces. Systematics After * †'' Umenocoleus'' Chen and Tan 1973 Dalazi Formation, China, Early Cretaceous (Aptian) Zhonggou Formation, China, Early Cretaceous (Albian) * †'' Ponopterix'' Vršanský and Grimaldi 1999 Crato Formation, Brazil, Early Cretaceous (Aptian) * †'' Blattapterix'' Vršanský 2003 Xiagou Formation, China, Early Cretaceous (Aptian) * †'' Elytropterix'' Vršanský 2003 Dzun-Bain Formation, Mongolia, Aptian * †'' Petropterix'' Vršanský 2003 Dzun-Bain Formation, Mongolia, Zaza Formation, Russia, Kitadani Formation, Japan, Early Cretaceous (Aptian) Zhonggou Formation, China, Early Cretaceous (Albian) * � ...
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Burmese Amber
Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The amber is of significant palaeontological interest due to the diversity of flora and fauna contained as inclusions, particularly arthropods including insects and arachnids but also birds, lizards, snakes, frogs and fragmentary dinosaur remains. The amber has been known and commercially exploited since the first century AD, and has been known to science since the mid-nineteenth century. Research on the deposit has attracted controversy due to its alleged role in funding internal conflict in Myanmar and hazardous working conditions in the mines where it is collected. Geological context, depositional environment and age The amber is found within the Hukawng Basin, a large Cretaceous- Cenozoic sedimentary basin within northern Myanmar. ...
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Aethiocarenus
''Aethiocarenus'' is an extinct genus of insects which has a single species ''Aethiocarenus burmanicus'' described from a 98.79 ±0.62 million year old fossil found in Burmese amber from the Hukawng Valley of Myanmar. The insect is unusual due to the vertex of the triangular head being attached to the pronotum as opposed to the hypotenuse. When first described ''Aethiocarenus'' was placed as the sole member of the family Aethiocarenidae and order Aethiocarenodea. However, ''Aethiocarenus'' was later considered to be a nymph of ''Alienopterus''. Vršanský ''et al.'' (2018) considered ''Aethiocarenus'' to be an alienopterid nymph, but considered it distinct from other members of this group and deserving a separate genus rank. Description ''Aethiocarenus'' was probably an omnivore An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, pro ...
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United States
The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five major unincorporated territories, nine United States Minor Outlying Islands, Minor Outlying Islands, and 326 Indian reservations. The United States is also in Compact of Free Association, free association with three Oceania, Pacific Island Sovereign state, sovereign states: the Federated States of Micronesia, the Marshall Islands, and the Palau, Republic of Palau. It is the world's List of countries and dependencies by area, third-largest country by both land and total area. It shares land borders Canada–United States border, with Canada to its north and Mexico–United States border, with Mexico to its south and has maritime borders with the Bahamas, Cuba, Russia, and other nations. With a population of over 333 million, it is the List of ...
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Colorado
Colorado (, other variants) is a state in the Mountain states, Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of the Great Plains. Colorado is the List of U.S. states and territories by area, eighth most extensive and List of U.S. states and territories by population, 21st most populous U.S. state. The 2020 United States Census, 2020 United States census enumerated the population of Colorado at 5,773,714, an increase of 14.80% since the 2010 United States Census, 2010 United States census. The region has been inhabited by Indigenous peoples of the Americas, Native Americans and their Paleo-Indians, ancestors for at least 13,500 years and possibly much longer. The eastern edge of the Rocky Mountains was a major migration route for early peoples who spread throughout the Americas. "''Colorado''" is the Spanish adjective meaning "ruddy", th ...
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Green River Formation
The Green River Formation is an Eocene geologic formation that records the sedimentation in a group of intermountain lakes in three basins along the present-day Green River in Colorado, Wyoming, and Utah. The sediments are deposited in very fine layers, a dark layer during the growing season and a light-hue inorganic layer in the dry season. Each pair of layers is called a varve and represents one year. The sediments of the Green River Formation present a continuous record of six million years. The mean thickness of a varve here is 0.18 mm, with a minimum thickness of 0.014 mm and maximum of 9.8 mm.Bradley, W. H. The varves and climate of the Green River epoch: U.S. Geol. Survey Prof. Paper 158, pp 87–110, 1929. The sedimentary layers were formed in a large area named for the Green River, a tributary of the Colorado River. The three separate basins lie around the Uinta Mountains (north, east, and south) of northeastern Utah: * an area in northwestern Colorado ea ...
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Brazil
Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area and the seventh most populous. Its capital is Brasília, and its most populous city is São Paulo. The federation is composed of the union of the 26 states and the Federal District. It is the largest country to have Portuguese as an official language and the only one in the Americas; one of the most multicultural and ethnically diverse nations, due to over a century of mass immigration from around the world; and the most populous Roman Catholic-majority country. Bounded by the Atlantic Ocean on the east, Brazil has a coastline of . It borders all other countries and territories in South America except Ecuador and Chile and covers roughly half of the continent's land area. Its Amazon basin includes a vast tropical forest, ho ...
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Crato Formation
The Crato Formation is a geologic formation of Early Cretaceous (Aptian) age in northeastern Brazil's Araripe Basin. It is an important Lagerstätte (undisturbed fossil accumulation) for palaeontologists. The strata were laid down mostly during the Aptian age, about 113 million years ago. It thought to have been deposited in a semi-arid lacustrine wetland environment.Ribeiro et al., 2021 The Crato Formation earns the designation of Lagerstätte due to an exceedingly well preserved and diverse fossil faunal assemblage. Some 25 species of fossil fishes are often found with stomach contents preserved, enabling paleontologists to study predator-prey relationships in this ecosystem. There are also fine examples of pterosaurs, reptiles and amphibians, invertebrates (particularly insects), and plants. Even dinosaurs are represented: a new maniraptor was described in 1996. The unusual taphonomy of the site resulted in limestone accretions that formed nodules around dead organisms, pre ...
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