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Admetophoneus
''Admetophoneus'' is a dubious genus of non-mammalian synapsid from Russia. Its type and only species is ''Admetophoneus kargalensis''. History ''Admetophoneus'' was named by the Russian paleontologist Ivan Efremov in 1954, based on some teeth, a fragmentary maxilla, and a humerus. It was originally classified as a member of Brithopodidae. Later, it was classified in Phthinosuchidae. Recent study, however, has shown that it is a member of Anteosauria, but lacks diagnostic features. It is essentially identical to the contemporaneous ''Titanophoneus'', and could be synonymous with it, but its poor preservation means that it cannot be proven. See also * List of therapsids This list of therapsids is an attempt to create a comprehensive listing of all Genus, genera that have ever been included in the Therapsida excluding mammals and purely vernacular terms. The list includes all commonly accepted genera, but also gene ... References The main groups of non-mammalian synapsids ...
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Anteosauria
Anteosaurs are a group of large, primitive carnivore, carnivorous dinocephalian therapsids with large canines and incisors and short limbs, that are known from the Middle Permian of South Africa, Russia, China, and Brazil. Some grew very large, with skulls long, and were the largest predators of their time. They died out at the end of the Middle Permian, possibly as a result of the extinction of the herbivorous Tapinocephalia on which they may have fed. Description The Anteosauria are distinguished from the Tapinocephalia by several features, such as very large canine (tooth), canines, cheek teeth with bulbous tooth crown, crowns, and an upturning of the premaxilla, so that the front of mouth curves strongly upwards. There is a tendency especially in more advanced forms such as ''Anteosaurus'' towards thickening of the bones of the top of the skull, indicating head-butting behavior. There is a large canal for the pineal organ (third eye); probably tied in with the animal's diurn ...
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List Of Therapsids
This list of therapsids is an attempt to create a comprehensive listing of all Genus, genera that have ever been included in the Therapsida excluding mammals and purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful (''nomen dubium, nomina dubia''), or were not formally published (''nomen nudum, nomina nuda''), as well as synonym (zoology), junior synonyms of more established names, and genera that are no longer considered therapsids. The list currently contains 510 generic names. Naming conventions and terminology Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include: * Synonym (zoology), Junior synonym: A name which describes the same taxon as a previously published name. If two or more genera are formally designated and the type (zoology), type specimens are later assigned to the same genus, the first to be published (in chronological o ...
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Anteosaurs
Anteosaurs are a group of large, primitive carnivorous dinocephalian therapsids with large canines and incisors and short limbs, that are known from the Middle Permian of South Africa, Russia, China, and Brazil. Some grew very large, with skulls long, and were the largest predators of their time. They died out at the end of the Middle Permian, possibly as a result of the extinction of the herbivorous Tapinocephalia on which they may have fed. Description The Anteosauria are distinguished from the Tapinocephalia by several features, such as very large canines, cheek teeth with bulbous crowns, and an upturning of the premaxilla, so that the front of mouth curves strongly upwards. There is a tendency especially in more advanced forms such as '' Anteosaurus'' towards thickening of the bones of the top of the skull, indicating head-butting behavior. There is a large canal for the pineal organ (third eye); probably tied in with the animal's diurnal and seasonal cycles. The shoul ...
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Guadalupian
The Guadalupian is the second and middle Series (stratigraphy), series/Epoch (geology), epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant Extinction event, mass extinction called the end-Capitanian extinction event. The Guadalupian is also known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was p ...
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Ivan Yefremov
Ivan Antonovich (Antipovich) Yefremov, sometimes Efremov (; 23 April 1908 – 5 October 1972) was a Soviet paleontologist, science-fiction author and social thinker. He founded taphonomy, the study of fossilization patterns. Biography He was born in the village of Vyritsa in Saint Petersburg Governorate on 23 April 1908. His parents divorced during the Russian Revolution. His mother married a Red Army commander and left the children in Kherson to be cared for by an aunt who soon died of typhus. Yefremov survived on his own for some time, after which he joined a Red Army unit as a "son of the regiment" and went to Perekop with it. In 1921, he was discharged and went to Petrograd (now Saint Petersburg) to study. He completed his education there while combining his studies with a variety of odd jobs. He later commented that "the Revolution was also my own liberation from philistinism" (). Academic career In 1956, due to the influence of academician Petr Sushkin, he bec ...
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Genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants of an ancestral taxon are grouped together (i.e. Phylogeneti ...
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Mammal
A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three Evolution of mammalian auditory ossicles, middle ear bones. These characteristics distinguish them from reptiles and birds, from which their ancestors Genetic divergence, diverged in the Carboniferous Period over 300 million years ago. Around 6,640 Neontology#Extant taxon, extant species of mammals have been described and divided into 27 Order (biology), orders. The study of mammals is called mammalogy. The largest orders of mammals, by number of species, are the rodents, bats, and eulipotyphlans (including hedgehogs, Mole (animal), moles and shrews). The next three are the primates (including humans, monkeys and lemurs), the Artiodactyl, even-toed ungulates (including pigs, camels, and whales), and the Carnivora (including Felidae, ...
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Synapsid
Synapsida is a diverse group of tetrapod vertebrates that includes all mammals and their extinct relatives. It is one of the two major clades of the group Amniota, the other being the more diverse group Sauropsida (which includes all extant reptiles and therefore, birds). Unlike other amniotes, synapsids have a single temporal fenestra, an opening low in the skull roof behind each eye socket, leaving a zygomatic arch, bony arch beneath each; this accounts for the name "synapsid". The distinctive temporal fenestra developed about 318 million years ago during the Late Carboniferous period, when synapsids and sauropsids diverged, but was subsequently merged with the orbit in early mammals. The basal (phylogenetics), basal amniotes (reptiliomorphs) from which synapsids evolved were historically simply called "reptiles". Therefore, stem group synapsids were then described as mammal-like reptiles in classical systematics, and non-therapsid synapsids were also referred to as pelyco ...
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Brithopodidae
''Brithopus'' is an extinct genus of dinocephalian therapsids. It contains a single species, ''Brithopus priscus'', known from fragmentary remains found in the Copper Sandstones near Isheevo, Russia. Description ''Brithopus'' was fairly large, reaching a length of 2.5–3 m (8–10 ft). The skull was similar to ''Titanophoneus'', but more massive and heavily built.Olson, E.C. (1962). "Lopingian, Late Permian terrestrial vertebrates, U.S.A. and U.S.S.R." ''Transactions of the American Philosophical Society'', New Series, 52: 1–224. Classification ''B. priscus'' was first named in 1838 and was traditionally classified in the Anteosauria, a group of carnivorous dinocephalians. ''Brithopus'' served as the basis for the family Brithopodidae, which once included many anteosaurian species. Because it is based on fragmentary material, ''Brithopus'' is regarded as a ''nomen dubium'' by some researchers. ''Brithopus'' was later considered a possible estemmenosuchid,Kammerer, C. F. 2 ...
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Phthinosuchidae
''Phthinosuchus'' is an extinct genus of therapsids from the Middle Permian of Russia. ''Phthinosuchus'' is the sole member of the family Phthinosuchidae. ''Phthinosuchus'' may have been one of the most primitive therapsids, meaning that its ancestors may have branched off early from the main therapsid line. Discovery ''Phthinosuchus'' was named in 1954 by Ivan Yefremov. It is currently only known from the back of the skull, as the front of the skull was lost after description. Two species, ''P. discors'' and ''P. horissiaki'', have been described. Description ''Phthinosuchus'' was 1.5 m (5 ft) long with a 20 cm skull, and looked much like the Sphenacodontidae, Sphenacodontids, such as ''Dimetrodon'' and ''Sphenacodon''. Its temporal fenestrae were larger than those of the Sphenacodontids. Its jaw was slender, unlike other predatory therapsids, but like the other Biarmosuchia, early therapsids, it was probably sprawling and carnivorous. Classification ''Phthinosu ...
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Titanophoneus
''Titanophoneus'' ("titanic murderer") is an extinct genus of carnivorous dinocephalian therapsid from the Middle Permian. It is classified within the family Anteosauridae. The type species is ''Titanophoneus potens''. Remains of ''Titanophoneus'' have been found at Isheevo, Russia. Description An adult skull would have reached 80 cm with a heavy long snout. The long tail and short limbs show the species to be a primitive therapsid, unlike '' Inostrancevia,'' which was more advanced. The structure of the limbs and the density of the bone are designed for a sprawling stance. The temporal opening is more advanced than Estemmenosuchidae, but less advanced than ''Inostrancevia''. The teeth are large with 12 large palate incisors followed by 2 canines and various smaller back teeth. The lower palate is the same as the upper but without the canine teeth. The appearance of ''Titanophoneus'' is reminiscent to the sphenacodontid pelycosaurs, which included '' Dimetrodon''. Ga ...
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Dinocephalian Genera
Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephalians included herbivorous, carnivorous, and omnivorous forms. Many species had thickened skulls with many knobs and bony projections. Dinocephalians were the first non-mammalian therapsids to be scientifically described and their fossils are known from Russia, China, Brazil, South Africa, Zimbabwe, and Tanzania. Description Apart from the biarmosuchians, the dinocephalians are the least advanced therapsids, although still uniquely specialised in their own way. They retain a number of primitive characteristics (e.g. no secondary palate, small dentary) shared with their pelycosaur ancestors, although they are also more advanced in possessing therapsid adaptations like the expansion of the ilium and more erect limbs. They include ...
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