''Sinornithoides'' (meaning "Chinese bird form") is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

of troodontid

theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

s containing the single

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...

''Sinornithoides youngi''. ''S. youngi'' lived during the

Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145& ...

(

Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ag ...

Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/ Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0 ...

stage Stage or stages may refer to: Acting * Stage (theatre), a space for the performance of theatrical productions * Theatre, a branch of the performing arts, often referred to as "the stage" * ''The Stage'', a weekly British theatre newspaper * Sta ...

, around 113

million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago ...

).Sereno, P.C. (2010). "Taxonomy, cranial morphology, and relationships of parrot-beaked dinosaurs (Ceratopsia: ''Psittacosaurus'')." ''New Perspectives on Horned Dinosaurs''. Bloomington: Indiana, 21-58. It measured approximately one

meter The metre (British spelling) or meter (American spelling; see spelling differences) (from the French unit , from the Greek noun , "measure"), symbol m, is the primary unit of length in the International System of Units (SI), though its pref ...

long (3.3 ft). It lived in

Inner Mongolia Inner Mongolia, officially the Inner Mongolia Autonomous Region, is an autonomous region of the People's Republic of China. Its border includes most of the length of China's border with the country of Mongolia. Inner Mongolia also accounts for a ...

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...

, and probably ate invertebrates and other small prey. They lived in what is now Mongolia, which was part of Laurasia.

Discovery

In 1988, a Chinese-Canadian expedition discovered the remains of a small theropod near Huamuxiao, in the Ordos Basin of Inner Mongolia. ''Sinornithoides youngi'', the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specim ...

, was named and described in 1993/1994 by Dale Russell and

Dong Zhiming Dong Zhiming (Chinese: 董枝明, Pinyin: ''Dǒng Zhimíng''; born January 1937) is a Chinese vertebrate paleontologist formerly employed at the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) in Beijing. He began working at th ...

based on this fossil specimen from the Lower Cretaceous

Ejinhoro Formation The Ejinhoro Formation () is a geological formation in Inner Mongolia, north China, whose strata date back to the Early Cretaceous period (Aptian/Albian age.Sereno, P.C. (2010). "Taxonomy, cranial morphology, and relationships of parrot-beaked d ...

. The generic name is derived from

Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...

''Sinae'', "Chinese", and Greek ὄρνις, ''ornis'', "bird", en ~ειδής, ''~eides'', a suffix meaning "~like", in reference to the bird-like build. The specific name honours

Yang Zhongjian Yang Zhongjian, also Yang Chung-chien (; 1 June 1897 – 15 January 1979), courtesy name Keqiang (), also known as C.C. (Chung Chien) Young, was a Chinese paleontologist and zoologist. He was one of China's foremost vertebrate paleontologists. H ...

.Russell, D. and Dong, Z. (1993). "A nearly complete skeleton of a new troodontid dinosaur from the Early Cretaceous of the Ordos Basin, Inner Mongolia, People's Republic of China." ''Canadian Journal of Earth Sciences'', 30: 2163-2173. doi:10.1139/e93-187 It is represented by a

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

, IVPP V9612, an almost complete skeleton with skull, which is articulated and nearly complete except for the roof of the skull, some cervical and many dorsal vertebrae, along with some other referred skeletal elements. The holotype is preserved in much the same roosting position as another troodontid fossil, that of '' Mei long'', with its snout tucked under its left hand. It represents a subadult individual.Currie, P. and Dong, Z. (2001). "New information on Cretaceous troodontids from the People's Republic of China." ''Canadian Journal of Earth Sciences'', 38: 1753–1766.

Description

''Sinornithoides'' is a

troodontid Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil disco ...

, a group of small, bird-like,

gracile Gracility is slenderness, the condition of being gracile, which means slender. It derives from the Latin adjective ''gracilis'' ( masculine or feminine), or ''gracile'' ( neuter), which in either form means slender, and when transferred for exa ...

maniraptorans. All troodontids have many unique features of the skull, such as closely spaced

teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, ...

in the lower jaw, and large numbers of teeth. Troodontids have sickle-claws and

raptorial The term ''raptorial'' implies much the same as ''predatory'' but most often refers to modifications of an arthropod's foreleg that make it function for the grasping of prey while it is consumed, where the gripping surfaces are formed from the o ...

hands A hand is a prehensile, multi- fingered appendage located at the end of the forearm or forelimb of primates such as humans, chimpanzees, monkeys, and lemurs. A few other vertebrates such as the koala (which has two opposable thumbs on eac ...

, and some of the highest non-

avian Avian may refer to: * Birds or Aves, winged animals *Avian (given name) (russian: Авиа́н, link=no), a male forename Aviation *Avro Avian, a series of light aircraft made by Avro in the 1920s and 1930s *Avian Limited, a hang glider manufactur ...

encephalization quotient Encephalization quotient (EQ), encephalization level (EL), or just encephalization is a relative brain size measure that is defined as the ratio between observed to predicted brain mass for an animal of a given size, based on nonlinear regress ...

s, meaning they were behaviourally advanced and had keen senses. In 2010, Gregory S. Paul estimated its body length at 1.1 metres, its weight at 2.5 kilogrammes.Paul, G.S., 2010, ''The Princeton Field Guide to Dinosaurs'', Princeton University Press p. 139 Sinornithoides-youngi_jconway

In 1994, ''Sinornithoides'' was one of the most completely known troodontids, especially as regarded the palate, but the lack of sufficient fossil material to compare it with induced Russell and Dong not to indicate any diagnostic traits. The skull of ''Sinornithoides'' is elongated and pointed. However, the head is relatively short compared to the body as a whole. The

praemaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has ...

is short. There is a ''fenestra promaxillaris'', a small opening in the front of the

maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. T ...

side, which is rare among troodontids. There are four premaxillary and about twenty-three maxillary teeth. The maxillary teeth have no serrations on their front edge; the denticles on the concavely curved rear edge are small. The maxillary teeth are rather recurved. The

lacrimal bone The lacrimal bone is a small and fragile bone of the facial skeleton; it is roughly the size of the little fingernail. It is situated at the front part of the medial wall of the orbit. It has two surfaces and four borders. Several bony landmarks of ...

lacks a pneumatic channel. In the braincase the subotic recess is large. The tips of the lower jaws do not curve towards each other but touch at their inner sides. The external mandibular fenestra is large. The dentary, lower jaw, teeth are quite pointed and have no front edge denticles; their rear edge is very straight. The holotype preserves a

furcula The (Latin for "little fork") or wishbone is a forked bone found in most birds and some species of non-avian dinosaurs, and is formed by the fusion of the two pink clavicles. In birds, its primary function is in the strengthening of the thoracic ...

and a basket of fifteen pairs of

gastralia Gastralia (singular gastralium) are dermal bones found in the ventral body wall of modern crocodilians and tuatara, and many prehistoric tetrapods. They are found between the sternum and pelvis, and do not articulate with the vertebrae. In thes ...

. The arms are weakly developed, with a slender

humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a r ...

and

ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...

. The front top of the third

metatarsal The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the me ...

is not fully overgrown by the second and fourth metatarsals. The sickle claw of the second toe is relatively large and long for a troodontid.

Classification

''Sinornithoides'' was in 1994 assigned to the

Troodontidae Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil discov ...

. A possible position of ''Sinornithoides'' in the evolutionary tree of the

Paraves Paraves are a widespread group of theropod dinosaurs that originated in the Middle Jurassic period. In addition to the extinct dromaeosaurids, troodontids, anchiornithids, and possibly the scansoriopterygids, the group also contains the avial ...

is shown by the

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...

below, following a 2012 analysis by Turner, Makovicky and Norell. Troodontid teeth

References

{{Taxonbar, from=Q134725 Early Cretaceous dinosaurs of Asia Troodontids Fossil taxa described in 1993 Taxa named by Dale Russell Taxa named by Dong Zhiming Paleontology in Inner Mongolia

Discovery

Description

Classification

See also

References