''Kapes'' is an extinct

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

procolophonid Procolophonidae is an extinct family of small, lizard-like parareptiles known from the Late Permian to Late Triassic that were distributed across Pangaea, having been reported from Europe, North America, China, South Africa, South America, Antarc ...

parareptile Parareptilia ("at the side of reptiles") is a subclass or clade of basal sauropsids (reptiles), typically considered the sister taxon to Eureptilia (the group that likely contains all living reptiles and birds). Parareptiles first arose near th ...

from the

Lower Lower may refer to: *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Gloucestershire, England See also *Nizhny Nizhny (russian: Ни́жний; masculine), Nizhnyaya (; feminine), or Nizhneye (russian: Ни́� ...

and

Middle Triassic In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between Ma an ...

of the United Kingdom and Russia. It is a member of the subfamily

Procolophoninae Procolophoninae is an extinct subfamily of procolophonid parareptiles from the late Early Triassic to the early Middle Triassic (Olenekian and Anisian stages) of Africa, Antarctica, Asia, Europe and South America. Currently, the oldest-known pr ...

. The type species ''K. amaenus'' was named in 1975 from the banks of the Vychegda River in the

Komi Republic The Komi Republic (russian: Республика Коми; kv, Коми Республика), sometimes simply referred to as Komi, is a republic of Russia located in Eastern Europe. Its capital is the city of Syktyvkar. The population of th ...

of Russia. In 1983, a new species was brought into the genus, ''K. majmesculae''. ''K. majmesculae'' was first named in 1968 as a member of the genus ''

Tichvinskia ''Tichvinskia'' is an extinct genus of procolophonid parareptile from the Early Triassic The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between Ma and Ma (million years ag ...

''. A third Russian species, ''K. serotinus'', was named in 1991. In 2002, ''Kapes bentoni'' was named from the

Otter Sandstone Formation The Otter Sandstone Formation is an obsolete geological unit in England, now referred to as the Helsby Sandstone Formation.http://www.bgs.ac.uk/Lexicon/lexicon.cfm?pub=HEY (BGS on-line lexicon of rock units) It preserves fossils dated to the Tr ...

Devon, England Devon ( , historically known as Devonshire , ) is a ceremonial and non-metropolitan county in South West England. The most populous settlement in Devon is the city of Plymouth, followed by Devon's county town, the city of Exeter. Devon is a ...

, extending the geographic range of ''Kapes''. In the same paper, ''K. serotinus'' was synonymized with ''K. majmesculae'' and another Russian species was assigned to ''Kapes'' called ''K. komiensis''. ''K. komiensis'' was first named in 1975 (in the same paper ''K. amaenus'' was named in) as a member of the genus '' Macrophon''.

Phylogeny

Below is a

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...

from Ruta ''et al.'' (2011) showing the placement of ''Kapes'' within Procolophonidae:

References

Procolophonines Triassic parareptiles Extinct animals of Europe Early Triassic genus first appearances Middle Triassic genus extinctions Prehistoric reptile genera {{paleo-reptile-stub