Amphisbaenidae
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The Amphisbaenidae (
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...
: worm lizards) are a
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of amphisbaenians, a group of limbless vertebrates.


Geographic range

Amphisbaenids occur in
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sou ...
, some
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean ...
islands,
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
, and
sub-Saharan Africa Sub-Saharan Africa is, geographically, the area and regions of the continent of Africa that lies south of the Sahara. These include West Africa, East Africa, Central Africa, and Southern Africa. Geopolitically, in addition to the List of sov ...
.


Taxonomy

One deep-branching and somewhat aberrant
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
, '' Blanus'', is native to Europe, and may represent a distinct family. More recent sources indeed place it in the family Blanidae.


Description

Members of the family Amphisbaenidae are limbless,
burrowing An Eastern chipmunk at the entrance of its burrow A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of locomotion. Burrows provide a form of s ...
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates ( lizards and snakes) and rhynchocephalia ...
s with
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other ...
diets. As in other amphisbaenians, the body bears rings of scales, which gives amphisbaenids a worm-like appearance. The head is massively constructed and used for burrowing, with powerful jaws and large, recurved teeth used for seizing prey. Some
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
have a
spade A spade is a tool primarily for digging consisting of a long handle and blade, typically with the blade narrower and flatter than the common shovel. Early spades were made of riven wood or of animal bones (often shoulder blades). After the a ...
-like head, while others have a narrow keel on the head, and still others have a rounded skull. The eyes are highly reduced, while the ear bone, or stapes in the
middle ear The middle ear is the portion of the ear medial to the eardrum, and distal to the oval window of the cochlea (of the inner ear). The mammalian middle ear contains three ossicles, which transfer the vibrations of the eardrum into waves in the ...
, is large and massive. Together with another bone, the extracollumella, the stapes detects vibrations caused by prey items, allowing amphisbaenids to hunt for invertebrates under ground. In this respect, apparently evolution exists convergent to the burrowing mammalian family
Chrysochloridae Golden moles are small insectivorous burrowing mammals endemic to Sub-Saharan Africa. They comprise the family Chrysochloridae and as such they are taxonomically distinct from the true moles, family Talpidae, and other mole-like families, all ...
, in which the malleus in the middle ear is greatly enlarged.


Extant genera

Over 170 extant species are in the family, grouped into 12
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
:


Fossil genera

A number of extinct taxa are known from the fossil record:


References


Further reading

* Gans C (2005). "Checklist and Bibliography of the Amphisbaenia of the World". ''Bulletin of the American Museum of Natural History'' (289): 1-130. * Goin CJ, Goin OB, Zug GR (1978). ''Introduction to Herpetology, Third Edition''. San Francisco: W.H. Freeman and Company. xi + 378 pp. . (Family Amphisbaenidae, pp. 276–277).


External links

*http://www.jcvi.org/reptiles/families/amphisbaenidae.php {{Taxonbar, from=Q733407 Amphisbaenians Reptile families Taxa named by John Edward Gray Extant Maastrichtian first appearances