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Paraneoptera
Paraneoptera or Acercaria is a superorder of insects which includes lice (bark lice and true lice), thrips, and hemipterans, the true bugs. It also includes the extinct order Permopsocida, known from fossils dating from the Early Permian to the mid-Cretaceous. All of the insects classified here exhibit various “reductions” or “simplifications” from the primitive body-plan found in typical polyneopterans. Cerci, for example, are entirely absent in all living paraneopterans (Acercaria meaning without cerci). Other “reductions” occur in wing venation, in the number of tarsal segments (no more than three), only four Malpighian tubules, and only one complex of abdominal ganglia. The mouthparts of the Paraneoptera reflect diverse feeding habits. Basal groups are microbial surface feeders, whereas more advanced groups feed on plant or animal fluids. Phylogeny Paraneoptera consists of Psocodea (lice), along with their sister clade, the monophyletic grouping Condylognat ...
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Psocodea
Psocodea is a taxonomic group of insects comprising the bark lice, book lice and parasitic lice. It was formerly considered a superorder, but is now generally considered by entomologists as an order. Despite the greatly differing appearance of parasitic lice (Phthiraptera), they are believed to have evolved from within the former order Psocoptera, which contained the bark lice and book lice, now found to be paraphyletic. They are often regarded as the most primitive of the hemipteroids. Psocodea contains around 11,000 species, divided among four suborders and more than 70 families. They range in size from 1–10 millimetres (0.04–0.4 in) in length. The species known as booklice received their common name because they are commonly found amongst old books—they feed upon the paste used in binding. The barklice are found on trees, feeding on algae and lichen. Anatomy and biology Psocids are small, scavenging insects with a relatively generalized body plan. They feed ...
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Permopsocida
Permopsocida is an extinct order of insects known from the Early Permian to the Mid-Cretaceous. It is part of Paraneoptera, alongside bark lice (including lice), bugs and thrips. Within Paraneoptera it is considered to be closer to the clade containing bugs and thrips rather than bark lice, with an estimated divergence during the Late Carboniferous. The group was first named as a suborder by Robert John Tillyard in 1926, and was raised to a full order by Huang et al. in 2016. It is currently divided up into three families, Psocidiidae which is known from the Permian to Liassic. Permopsocidae which is only known from the Permian, and Archipsyllidae, which is known from the Late Triassic to mid-Cretaceous (Cenomanian). While most members of the group are known from compression fossils, several members of Archipsyllidae are 3 dimensionally preserved in Burmese amber, which has helped clarify the morphology and phylogenetic position of the group. The morphology of the mouthparts sugg ...
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Hemiptera
Hemiptera (; ) is an order of insects, commonly called true bugs, comprising over 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, assassin bugs, bed bugs, and shield bugs. They range in size from to around , and share a common arrangement of piercing-sucking mouthparts. The name "true bugs" is often limited to the suborder Heteroptera. Entomologists reserve the term ''bug'' for Hemiptera or Heteroptera,Gilbert Waldbauer. ''The Handy Bug Answer Book.'' Visible Ink, 1998p. 1. which does not include other arthropods or insects of other orders such as ants, bees, beetles, or butterflies. In some variations of English, all terrestrial arthropods (including non-insect arachnids, and myriapods) also fall under the colloquial understanding of ''bug''. Many insects with "bug" in their common name, especially in American English, belong to other orders; for example, the lovebug is a fly and the Maybug and ladybug are beetles. The term is al ...
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Hemiptera
Hemiptera (; ) is an order of insects, commonly called true bugs, comprising over 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, assassin bugs, bed bugs, and shield bugs. They range in size from to around , and share a common arrangement of piercing-sucking mouthparts. The name "true bugs" is often limited to the suborder Heteroptera. Entomologists reserve the term ''bug'' for Hemiptera or Heteroptera,Gilbert Waldbauer. ''The Handy Bug Answer Book.'' Visible Ink, 1998p. 1. which does not include other arthropods or insects of other orders such as ants, bees, beetles, or butterflies. In some variations of English, all terrestrial arthropods (including non-insect arachnids, and myriapods) also fall under the colloquial understanding of ''bug''. Many insects with "bug" in their common name, especially in American English, belong to other orders; for example, the lovebug is a fly and the Maybug and ladybug are beetles. The term is al ...
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Neoptera
Neoptera (Ancient Greek ''néos'' (“new”) + ''pterón'' (“wing”)) is a classification group that includes most orders of the winged insects, specifically those that can flex their wings over their abdomens. This is in contrast with the more basal orders of winged insects (the " Palaeoptera" assemblage), which are unable to flex their wings in this way. Classification The taxon Neoptera was proposed by А.М. Martynov in 1923 and 1924, in the following classification: Pterygota *division Palaeoptera **order Odonata **order Agnatha (correct name: Ephemeroptera) **†order Dictyoneuridea **†order Megasecoptera **†order Meganisoptera **†order Protephemeroidea *division Neoptera **superorder Endopterygota **subdivision Polyneoptera ***superorder Orthopteroidea (Anartioptera) ****order Orthoptera ****order Plecoptera ****order Dermaptera ****order Embioptera ****order Phasmatodea ***superorder Blattopteroidea (senior name: Pandictyoptera) ****order Blattodea *** ...
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Insect
Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs of jointed legs, compound eyes and one pair of antennae. Their blood is not totally contained in vessels; some circulates in an open cavity known as the haemocoel. Insects are the most diverse group of animals; they include more than a million described species and represent more than half of all known living organisms. The total number of extant species is estimated at between six and ten million; In: potentially over 90% of the animal life forms on Earth are insects. Insects may be found in nearly all environments, although only a small number of species reside in the oceans, which are dominated by another arthropod group, crustaceans, which recent research has indicated insects are nested within. Nearly all insects hatch f ...
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Condylognatha
Condylognatha or Panhemiptera is a monophyletic grouping (superorder) that contains Hemiptera (true bugs) and Thysanoptera (thrips). Condylognatha belongs to Paraneoptera, which include its sister group, lice (Psocodea). Taxonomy Hemiptera and Thysanoptera are both the sister-groups based on morphological characters, and jointly known as Condylognatha. Hemiptera Hemiptera is an order of insects most often known as the ''true bugs'' (''cf.'' bug), comprising around 50,000–80,000 species of cicadas, aphids, planthoppers, leafhoppers, shield bugs, bed bugs and others. They range in size from to around , and share a common arrangement of sucking mouthparts. Thrips Order Thysanoptera includes 5,500 species classified into two suborders distinguished by the ovipositor. Terebrantia have a well-developed conical ovipositor, while the Tubulifera Phlaeothripidae is a family of thrips with hundreds of genera. They are the only extant family of the suborder Tubulifera ...
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Eumetabola
Eumetabola is an unranked clade of Neoptera. Two large unities known as the Paurometabola and Eumetabola are probably from the adelphotaxa of the Neoptera after exclusion of the Plecoptera. The monophyly of these unities appears to be weakly justified. Eumetabola has the highest number of species of any clade. According to a phylogenetic analysis, the Eumetabola clade originated 390-350 million years ago, in the Late Devonian.Wang, Y.-h. et al. Fossil record of stem groups employed in evaluating the chronogram of insects (Arthropoda: Hexapoda). Sci. Rep. 6, 38939; doi: 10.1038/srep38939 (2016). Phylogeny The phylogeny of Eumetabola is shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... according to Kluge 2004, 2010, and 2012: References Externa ...
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Polyneoptera
The cohort Polyneoptera is a proposed taxonomic ranking for the Orthoptera (grasshoppers, crickets, etc.) and all other Neopteran insects believed to be more closely related to Orthoptera than to any other insect orders. These winged insects, now in the Paraneoptera, were formerly grouped as the Hemimetabola or Exopterygota on the grounds that they have no metamorphosis, the wings gradually developing externally throughout the nymphal stages. Taxonomy The ''Polyneoptera Species File'' lists the following: Superorder Dermapterida * † Protelytroptera Superorder Dictyoptera * Blattodea – cockroaches and termites * Mantodea – praying mantises Superorder Orthopterida Synonyms include: Archaeorthoptera, Gryllidea, Orthopterodea, Orthopterodida, Orthopteroidea, Panorthoptera * † Caloneurodea * † Geraroptera * Orthoptera – 2 extant suborders: ** Caelifera – grasshoppers, groundhoppers, pygmy mole-crickets ** Ensifera – crickets, mole-crickets, katydids or ...
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Psocoptera
Psocoptera are a paraphyletic group of insects that are commonly known as booklice, barklice or barkflies. The name Psocoptera has been replaced with Psocodea in recent literature, with the inclusion of the former order Phthiraptera into Psocodea (as part of the suborder Troctomorpha). These insects first appeared in the Permian period, 295–248 million years ago. They are often regarded as the most primitive of the hemipteroids. Their name originates from the Greek word ψῶχος, ''psokhos'' meaning gnawed or rubbed and πτερά, ''ptera'' meaning wings. There are more than 5,500 species in 41 families in three suborders. Many of these species have only been described in recent years. They range in size from 1–10 millimetres (0.04–0.4 in) in length. The species known as booklice received their common name because they are commonly found amongst old books—they feed upon the paste used in binding. The barklice are found on trees, feeding on algae and ...
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Sister Clade
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and taxon B are sister groups to each other. Taxa A and B, together with any other extant or extinct descendants of their most recent common ancestor (MRCA), form a monophyletic group, the clade AB. Clade AB and taxon C are also sister groups. Taxa A, B, and C, together with all other descendants of their MRCA form the clade ABC. The whole clade ABC is itself a subtree of a larger tree which offers yet more sister group relationships, both among the leaves and among larger, more deeply rooted clades. The tree structure shown connects through its root to the rest of the universal tree of life. In cladistic standards, taxa A, B, and C may represent specimens, species, genera, or any other taxonomic units. If A and B are at the same taxo ...
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Phthiraptera
Louse ( : lice) is the common name for any member of the clade Phthiraptera, which contains nearly 5,000 species of wingless parasitic insects. Phthiraptera has variously been recognized as an order, infraorder, or a parvorder, as a result of developments in phylogenetic research. Lice are obligate parasites, living externally on warm-blooded hosts which include every species of bird and mammal, except for monotremes, pangolins, and bats. Lice are vectors of diseases such as typhus. Chewing lice live among the hairs or feathers of their host and feed on skin and debris, while sucking lice pierce the host's skin and feed on blood and other secretions. They usually spend their whole life on a single host, cementing their eggs, called nits, to hairs or feathers. The eggs hatch into nymphs, which moult three times before becoming fully grown, a process that takes about four weeks. Genetic evidence indicates that lice are a highly modified lineage of Psocoptera (now called Psoco ...
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