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Ascomycota
Ascomycota is a phylum of the kingdom Fungi that, together with the Basidiomycota, forms the subkingdom Dikarya. Its members are commonly known as the sac fungi or ascomycetes. It is the largest phylum of Fungi, with over 64,000 species. The defining feature of this fungal group is the " ascus" (), a microscopic sexual structure in which nonmotile spores, called ascospores, are formed. However, some species of the Ascomycota are asexual, meaning that they do not have a sexual cycle and thus do not form asci or ascospores. Familiar examples of sac fungi include morels, truffles, brewers' and bakers' yeast, dead man's fingers, and cup fungi. The fungal symbionts in the majority of lichens (loosely termed "ascolichens") such as ''Cladonia'' belong to the Ascomycota. Ascomycota is a monophyletic group (it contains all descendants of one common ancestor). Previously placed in the Deuteromycota along with asexual species from other fungal taxa, asexual (or anamorphic) ascomyce ...
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Fungi
A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from the other eukaryotic kingdoms, which by one traditional classification include Plantae, Animalia, Protozoa, and Chromista. A characteristic that places fungi in a different kingdom from plants, bacteria, and some protists is chitin in their cell walls. Fungi, like animals, are heterotrophs; they acquire their food by absorbing dissolved molecules, typically by secreting digestive enzymes into their environment. Fungi do not photosynthesize. Growth is their means of mobility, except for spores (a few of which are flagellated), which may travel through the air or water. Fungi are the principal decomposers in ecological systems. These and other differences place fungi in a single group of related organisms, named the ''Eumycota'' (''t ...
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Neolectomycetes
''Neolecta'' is a genus of ascomycetous fungi that have fruiting bodies in the shape of unbranched to lobed bright yellowish, orangish to pale yellow-green colored, club-shaped, smooth, fleshy columns up to about 7 cm tall. The species share the English designation "Earth tongues" along with some better-known fungi (e.g. ''Geoglossum'', ''Microglossum'') with a similar general form, but in fact they are only distantly related. ''Neolecta'' is the only genus belonging to the family Neolectaceae, which is the only family belonging to the order Neolectales. ''Neolectales'', in turn, is the only order belonging to the class Neolectomycetes, which belongs to the subdivision Taphrinomycotina of the Ascomycota. ''Neolecta'' is found in Asia, North America, Northern Europe and southern Brazil. The species all live in association with trees, and at least one, ''N. vitellina'', grows from rootlets of its host, but it is not known whether the fungus is parasitic, saprotrophic, or mutu ...
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Pezizomycotina
Pezizomycotina make up most of the Ascomycota fungi and include most lichenized fungi too. Pezizomycotina contains the filamentous ascomycetes and is a subdivision of the Ascomycota (fungi that form their spores in a sac-like ''ascus''). It is more or less synonymous with the older taxon Euascomycota. These fungi reproduce by fission rather than budding and this subdivision includes almost all the ascus fungi that have fruiting bodies visible to the naked eye (exception: genus ''Neolecta'', which belongs to the Taphrinomycotina). See the taxobox for a list of the classes that make up the Pezizomycotina. The old class Loculoascomycetes (consisting of all the bitunicate Ascomycota) has been replaced by the two classes Eurotiomycetes and Dothideomycetes. The rest of the Pezizomycotina also include the previously defined hymenial groups Discomycetes (now Leotiomycetes) and Pyrenomycetes (Sordariomycetes). Some important groups in Pezizomycotina include: Pezizomycetes (the opercula ...
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Saccharomycotina
Saccharomycotina is a subdivision (subphylum) of the division (phylum) Ascomycota in the kingdom Fungi. It comprises most of the ascomycete yeasts. The members of Saccharomycotina reproduce by budding and they do not produce ascocarps (fruiting bodies). The subdivision includes a single class: Saccharomycetes, which again contains a single order: Saccharomycetales. Notable members of Saccharomycotina are the baker's yeast ''Saccharomyces cerevisiae'' and the genus '' ''Candida'''' that includes several human pathogens. Etymology The name comes from the Greek word σάκχαρον (''sákkharon''), meaning "sugar" and μύκης (''mukēs'') meaning "fungus". History and economic importance Historical records from ancient Egypt and China describe the processes of brewing and baking from 10,000 to 8,000 years ago, and the production of fermented beverages and foods seems to have paralleled the beginning of agriculture. In the 1850s, Louis Pasteur demonstrated that yeasts a ...
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Taphrinomycotina
The Taphrinomycotina are one of three subdivisions constituting the Ascomycota (fungi that form their spores in a sac-like ascus) and is more or less synonymous with the slightly older invalid name Archiascomycetes (sometimes spelled Archaeascomycetes; archea = ancient). Recent molecular studies suggest that the group is monophyletic and basal to the rest of the Ascomycota. The major taxa are Schizosaccharomycetes, Taphrinomycetes, Neolectomycetes, and Pneumocystis. The Schizosaccharomycetes are the yeasts (e.g. ''Schizosaccharomyces'') that reproduce by fission rather than budding, unlike most other yeasts, many of which are in the subdivision Saccharomycotina. The Taphrinomycetes are dimorphic plant parasites (e.g. ''Taphrina'') with both a yeast state and a filamentous (hyphal) state in infected plants. They characteristically infect leaves, catkins, and branches, not roots. Taphrinomycetes form asci but no ascomata. The Neolectomycetes are species in a single genus, ''N ...
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Coniocybomycetes
The Coniocybaceae are the sole family of lichen-forming fungi in the Coniocybales, which itself is the only order in the class Coniocybomycetes. The family was circumscribed by Heinrich Gottlieb Ludwig Reichenbach in 1837. Both the order and the class were proposed by Maria Prieto and Mats Wedin in 2013 after molecular phylogenetics analysis of various calicioid lichens showed that the Coniocybaceae represented an early diverging lineage in the inoperculate ascomycete Ascomycota is a phylum of the kingdom Fungi that, together with the Basidiomycota, forms the subkingdom Dikarya. Its members are commonly known as the sac fungi or ascomycetes. It is the largest phylum of Fungi, with over 64,000 species. The defi ...s. Genera *'' Chaenotheca'' – 28 spp. *'' Coniocybe'' – 4 spp. *'' Sclerophora'' – 4 spp. References Ascomycota Ascomycota families Lichen families Taxa named by Ludwig Reichenbach Taxa described in 1837 {{Ascomycota-stub ...
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Geoglossomycetes
Geoglossaceae is a family of fungi in the order Geoglossales, class Geoglossomycetes. These fungi are broadly known as earth tongues. The ascocarps of most species in the family Geoglossaceae are terrestrial and are generally small, dark in color, and club-shaped with a height of 2–8 cm. The ascospores are typically light-brown to dark-brown and are often multiseptate. Other species of fungi have been known to parasitize ascocarps. The use of a compound microscope is needed for accurate identification."Family: Geoglossaceae." The Hidden Forest. 26 Aug. 2009 . Systematics The fungi that are now included in the fungal class Geoglossomycetes were previously considered by mycologists to be a family (Geoglossaceae) within the class Leotiomycetes. The family Geoglossaceae ''sensu lato'' was previously defined with 6 genera and 48 species. Early molecular evidence using ribosomal DNA suggested that Geoglossaceae ''sensu lato'' was not a monophyletic group, and that the hyaline s ...
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Lahmiales
The Lahmiales are an order of fungi in the Ascomycota, or sac fungi. The order has not been assigned to any class (''incertae sedis''). The taxon is monotypic and contains a single family, the Lahmiaceae, which in turn contains the single genus ''Lahmia''. The genus name of ''Lahmia'' is in honour of Johann Gottlieb Franz-Xaver Lahm (1811-1888), who was a German clergyman and botanist (Lichenology and Mycology). The genus of ''Lahmia'' was circumscribed In geometry, the circumscribed circle or circumcircle of a polygon is a circle that passes through all the vertices of the polygon. The center of this circle is called the circumcenter and its radius is called the circumradius. Not every polyg ... by Gustav Wilhelm Koerber in Parerga Lichenol on page 281 in 1861. References Pezizomycotina Ascomycota orders {{ascomycota-stub ...
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Sordariomycetes
Sordariomycetes is a class of fungi in the subdivision Pezizomycotina (Ascomycota), consisting of 28 orders, 90 families, 1344 genera. Sordariomycetes is from the Latin sordes (filth) because some species grow in animal feces, though growth habits vary widely across the class. Sordariomycetes generally produce their asci in perithecial fruiting bodies. Sordariomycetes are also known as Pyrenomycetes, from the Greek πυρἠν - 'the stone of a fruit' - because of the usually somewhat tough texture of their tissue. Sordariomycetes possess great variability in morphology, growth form, and habitat. Most have perithecial (flask-shaped) fruiting bodies, but ascomata can be less frequently cleistothecial (like in the genera '' Anixiella'', ''Apodus'', '' Boothiella'', ''Thielavia'', '' Zopfiella''),. Fruiting bodies may be solitary or gregarious, superficial, or immersed within stromata or tissues of the substrates and can be light to bright or black. Members of this group can grow ...
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Orbiliomycetes
The Orbiliaceae are a family of saprobic sac fungi in the order Orbiliales. The family, first described by John Axel Nannfeldt in 1932, contains 288 species in 12 genera. Members of this family have a widespread distribution, but are more prevalent in temperate regions. Some species in the Orbiliaceae are carnivorous fungi, and have evolved a number of specialized mechanisms to trap nematodes. Description Orbiliaceae do not have stromata, dense structural tissue that produces fruit bodies. They have small disc-shaped apothecia, that are typically convex, brightly colored or translucent. Their ascospores are small (typically less than 10 x 1 μm), hyaline, and have an oval or ellipsoidal shape. Species are usually found in wood on both wet and dry habitats. Anamorph species are hyphomycetous. Nematophagy This family is well known for its many nematophagous species. Shortly after coming into contact with its prey, fungal mycelia penetrate the nematode and spontaneously diffe ...
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Eurotiomycetes
Eurotiomycetes is a large class of ascomycetes with cleistothecial ascocarps within the subphylum Pezizomycotina, currently containing around 3810 species according to the Catalogue of Life. It is the third largest lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship. It contains most of the fungi previously known morphologically as "Plectomycetes".


Systematics and phylogeny


Internal relationships

The class Eurotiomycetes was circumscription (taxonomy), circumscribed in 1997 by Sweden, Swedish mycologists Ove Erik Eriksson and Katarina Winka. At that time ...

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Schizosaccharomycetes
Schizosaccharomycetes is a class in the kingdom of fungi. It contains the order Schizosaccharomycetales, the fission yeasts. The genus Schizosaccharomycetes is a broad and ancient clade within Ascomycota including five known fission yeast: ''Schizosaccharomyces pombe ''Schizosaccharomyces pombe'', also called "fission yeast", is a species of yeast used in traditional brewing and as a model organism in molecular and cell biology. It is a unicellular eukaryote, whose cells are rod-shaped. Cells typically meas ...'', ''Schizosaccharomyces japonicius'', ''Schizosaccharomyces octosporus'', and ''Schizosaccharomyces cryophilus'', and ''Schizosaccharomyces osmophilus''. References Yeasts Fungus classes Taxa described in 1997 {{yeast-stub ...
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