A midge is any small
fly
Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced ...
, including species in several
families of non-
mosquito Nematocera
The Nematocera (the name means "thread-horns") are a suborder of elongated flies with thin, segmented antennae and mostly aquatic larvae. This group is paraphyletic and contains all flies but species from suborder Brachycera (the name means "sh ...
n
Diptera. Midges are found (seasonally or otherwise) on practically every land area outside permanently arid deserts and the frigid zones. Some midges, such as many
Phlebotominae (sand fly) and
Simuliidae (black fly), are vectors of various diseases. Many others play useful roles as prey for
insectivores, such as various
frogs and
swallows. Others are important as
detritivores, and form part of various
nutrient cycles
A nutrient cycle (or ecological recycling) is the movement and exchange of inorganic and organic matter back into the production of matter. Energy flow is a unidirectional and noncyclic pathway, whereas the movement of mineral nutrients is cycli ...
. The habits of midges vary greatly from species to species, though within any particular family, midges commonly have similar ecological roles.
Examples of families that include species of midges include:
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Blephariceridae
The Blephariceridae, commonly known as net-winged midges, are a nematoceran family in the order Diptera. The adults resemble crane flies except with a projecting anal angle in the wings, and different head shape, absence of the V on the mesonotum, ...
, net-winged midges
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Cecidomyiidae, gall midges
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Ceratopogonidae, biting midges (also known as no-see-ums or punkies in
North America
North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
br>
BugGuide/ref> and sandflies in Australia)
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Chaoboridae
Chaoboridae, commonly known as phantom midges or glassworms, is a family of fairly common midges with a cosmopolitan distribution. They are closely related to the Corethrellidae and Chironomidae; the adults are differentiated through peculiaritie ...
, phantom midges
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Chironomidae, non-biting midges (also known as muckleheads, muffleheads or lake flies in the
Great Lakes
The Great Lakes, also called the Great Lakes of North America, are a series of large interconnected freshwater lakes in the mid-east region of North America that connect to the Atlantic Ocean via the Saint Lawrence River. There are five lakes ...
region of
North America
North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
)
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Deuterophlebiidae, mountain midges
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Dixidae, meniscus midges
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Scatopsidae, dung midges
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Thaumaleidae, solitary midges
Examples
The
Ceratopogonidae (biting midges) include serious blood-sucking pests, feeding both on humans and other mammals. Some of them spread the livestock diseases
blue tongue and
African horse sickness
African horse sickness (AHS) is a highly infectious and deadly disease caused by ''African horse sickness virus''. It commonly affects horses, mules, and donkeys. It is caused by a virus of the genus ''Orbivirus'' belonging to the family ' ...
– other species though, are at least partly nectar feeders, and some even suck insect bodily fluids.
Other ceratopogonid midges are major pollinators of ''
Theobroma cacao'' (cocoa tree). Having natural pollinators has beneficial effects in both agricultural and biological products because it increases crop yield and also density of predators of the midges (still beneficial to all parties).
Most other midge families are bloodsuckers, but it is not possible to generalise rigidly because of the vagueness of the term "midge". There is, for example, no objective basis for excluding the
Psychodidae from the list, and some of them (or midge-like taxa commonly included in the family, such as ''
Phlebotomus
''Phlebotomus'' is a genus of "sand flies" in the Diptera family Psychodidae. In the past, they have sometimes been considered to belong in a separate family, Phlebotomidae, but this alternative classification has not gained wide acceptance.
...
'') are blood-sucking pests and disease vectors.
Most midges, apart from the gall midges (
Cecidomyiidae), are aquatic during the larval stage. Some Cecidomyiidae (e.g., the sorghum midge) are significant plant pests. The larvae of some
Chironomidae contain
hemoglobin
Hemoglobin (haemoglobin BrE) (from the Greek word αἷμα, ''haîma'' 'blood' + Latin ''globus'' 'ball, sphere' + ''-in'') (), abbreviated Hb or Hgb, is the iron-containing oxygen-transport metalloprotein present in red blood cells (erythroc ...
and are sometimes referred to as
bloodworms.
Non-biting midge flies are a common minor nuisance around artificially-created bodies of water.
See also
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Highland midge
References
Further reading
{{cite journal, doi=10.1016/j.ecoenv.2020.111778, title=Impact of 2,4-D and fipronil on the tropical midge Chironomus sancticaroli (Diptera: Chironomidae), year=2021, last1=Pinto, first1=Thandy Junio da Silva, last2=Moreira, first2=Raquel Aparecida, last3=Silva, first3=Laís Conceição Menezes da, last4=Yoshii, first4=Maria Paula Cardoso, last5=Goulart, first5=Bianca Veloso, last6=Fraga, first6=Priscille Dreux, last7=Montagner, first7=Cassiana Carolina, last8=Daam, first8=Michiel Adriaan, last9=Espindola, first9=Evaldo Luiz Gaeta, journal=Ecotoxicology and Environmental Safety, volume=209, page=111778, pmid=33338803, doi-access=free
Nematocera
Insect common names