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Marine biology is the scientific study of the biology of marine life, organisms in the sea. Given that in biology many
phyla Phyla, the plural of ''phylum'', may refer to: * Phylum, a biological taxon between Kingdom and Class * by analogy, in linguistics, a large division of possibly related languages, or a major language family which is not subordinate to another Phy ...
, families and genera have some species that live in the sea and others that live on land, marine biology classifies species based on the
environment Environment most often refers to: __NOTOC__ * Natural environment, all living and non-living things occurring naturally * Biophysical environment, the physical and biological factors along with their chemical interactions that affect an organism or ...
rather than on taxonomy. A large proportion of all life on Earth lives in the ocean. The exact size of this ''large proportion'' is unknown, since many ocean species are still to be discovered. The ocean is a complex three-dimensional world covering approximately 71% of the Earth's surface. The habitats studied in marine biology include everything from the tiny layers of surface water in which organisms and abiotic items may be trapped in
surface tension Surface tension is the tendency of liquid surfaces at rest to shrink into the minimum surface area possible. Surface tension is what allows objects with a higher density than water such as razor blades and insects (e.g. water striders) t ...
between the ocean and atmosphere, to the depths of the oceanic trenches, sometimes 10,000 meters or more beneath the surface of the ocean. Specific habitats include
estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environme ...
,
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of Colony (biology), colonies of coral polyp (zoology), polyps held together by calcium carbonate. Most coral reefs are built from stony corals, wh ...
s, kelp forests, seagrass meadows, the surrounds of seamounts and thermal vents, tidepools, muddy, sandy and rocky bottoms, and the open ocean ( pelagic) zone, where solid objects are rare and the surface of the water is the only visible boundary. The organisms studied range from microscopic phytoplankton and zooplankton to huge cetaceans (whales) in length. Marine ecology is the study of how marine organisms interact with each other and the environment. Marine life is a vast resource, providing food, medicine, and raw materials, in addition to helping to support recreation and tourism all over the world. At a fundamental level, marine life helps determine the very nature of our planet. Marine organisms contribute significantly to the oxygen cycle, and are involved in the regulation of the Earth's
climate Climate is the long-term weather pattern in an area, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteorologi ...
. Shorelines are in part shaped and protected by marine life, and some marine organisms even help create new land. Many species are economically important to humans, including both finfish and shellfish. It is also becoming understood that the well-being of marine organisms and other organisms are linked in fundamental ways. The human body of knowledge regarding the relationship between life in the sea and important cycles is rapidly growing, with new discoveries being made nearly every day. These cycles include those of matter (such as the
carbon cycle The carbon cycle is the biogeochemical cycle by which carbon is exchanged among the biosphere, pedosphere, geosphere, hydrosphere, and atmosphere of the Earth. Carbon is the main component of biological compounds as well as a major compon ...
) and of air (such as Earth's respiration, and movement of energy through
ecosystem An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syst ...
s including the ocean). Large areas beneath the ocean surface still remain effectively unexplored.


Biological oceanography

Marine biology can be contrasted with biological oceanography. Marine life is a field of study both in marine biology and in biological oceanography. Biological oceanography is the study of how organisms affect and are affected by the
physics Physics is the natural science that studies matter, its fundamental constituents, its motion and behavior through space and time, and the related entities of energy and force. "Physical science is that department of knowledge which ...
,
chemistry Chemistry is the scientific study of the properties and behavior of matter. It is a natural science that covers the elements that make up matter to the compounds made of atoms, molecules and ions: their composition, structure, proper ...
, and
geology Geology () is a branch of natural science concerned with Earth and other Astronomical object, astronomical objects, the features or rock (geology), rocks of which it is composed, and the processes by which they change over time. Modern geology ...
of the oceanographic system. Biological oceanography mostly focuses on the
microorganism A microorganism, or microbe,, ''mikros'', "small") and ''organism'' from the el, ὀργανισμός, ''organismós'', "organism"). It is usually written as a single word but is sometimes hyphenated (''micro-organism''), especially in old ...
s within the ocean; looking at how they are affected by their environment and how that affects larger marine creatures and their ecosystem.Lalli, Carol M., and Timothy R. Parsons. "Introduction." Biological Oceanography: An Introduction. First Edition ed. Tarrytown, New York: Pergamon, 1993. 7-21. Print. Biological oceanography is similar to marine biology, but it studies ocean life from a different perspective. Biological oceanography takes a bottom up approach in terms of the food web, while marine biology studies the ocean from a top down perspective. Biological oceanography mainly focuses on the ecosystem of the ocean with an emphasis on plankton: their diversity (morphology, nutritional sources, motility, and metabolism); their productivity and how that plays a role in the global carbon cycle; and their distribution (predation and life cycle). Biological oceanography also investigates the role of microbes in food webs, and how humans impact the ecosystems in the oceans.


Marine habitats

Marine habitats can be divided into coastal and open ocean habitats. Coastal habitats are found in the area that extends from the shoreline to the edge of the continental shelf. Most marine life is found in coastal habitats, even though the shelf area occupies only seven percent of the total ocean area. Open ocean habitats are found in the deep ocean beyond the edge of the continental shelf. Alternatively, marine habitats can be divided into pelagic and demersal habitats. Pelagic habitats are found near the surface or in the open water column, away from the bottom of the ocean and affected by
ocean current An ocean current is a continuous, directed movement of sea water generated by a number of forces acting upon the water, including wind, the Coriolis effect, breaking waves, cabbeling, and temperature and salinity differences. Depth conto ...
s, while demersal habitats are near or on the bottom. Marine habitats can be modified by their inhabitants. Some marine organisms, like corals, kelp and sea grasses, are ecosystem engineers which reshape the marine environment to the point where they create further habitat for other organisms.


Intertidal and near shore

Intertidal zones, the areas that are close to the shore, are constantly being exposed and covered by the ocean's tides. A huge array of life can be found within this zone. Shore habitats span from the upper intertidal zones to the area where land vegetation takes prominence. It can be underwater anywhere from daily to very infrequently. Many species here are scavengers, living off of sea life that is washed up on the shore. Many land animals also make much use of the shore and intertidal habitats. A subgroup of organisms in this habitat bores and grinds exposed rock through the process of
bioerosion Bioerosion describes the breakdown of hard ocean substrates – and less often terrestrial substrates – by living organisms. Marine bioerosion can be caused by mollusks, polychaete worms, phoronids, sponges, crustaceans, echinoids, and ...
.


Estuaries

Estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environme ...
are also near shore and influenced by the tides. An estuary is a partially enclosed coastal body of water with one or more rivers or streams flowing into it and with a free connection to the open sea. Estuaries form a transition zone between freshwater river environments and saltwater maritime environments. They are subject both to marine influences—such as tides, waves, and the influx of saline water—and to riverine influences—such as flows of fresh water and sediment. The shifting flows of both sea water and fresh water provide high levels of nutrients both in the water column and in sediment, making estuaries among the most productive natural habitats in the world.


Reefs

Reefs comprise some of the densest and most diverse habitats in the world. The best-known types of reefs are tropical
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of Colony (biology), colonies of coral polyp (zoology), polyps held together by calcium carbonate. Most coral reefs are built from stony corals, wh ...
s which exist in most tropical waters; however, reefs can also exist in cold water. Reefs are built up by corals and other
calcium Calcium is a chemical element with the symbol Ca and atomic number 20. As an alkaline earth metal, calcium is a reactive metal that forms a dark oxide-nitride layer when exposed to air. Its physical and chemical properties are most similar ...
-depositing animals, usually on top of a rocky outcrop on the ocean floor. Reefs can also grow on other surfaces, which has made it possible to create artificial reefs. Coral reefs also support a huge community of life, including the corals themselves, their symbiotic zooxanthellae, tropical fish and many other organisms. Much attention in marine biology is focused on coral reefs and the El Niño weather phenomenon. In 1998, coral reefs experienced the most severe mass bleaching events on record, when vast expanses of reefs across the world died because sea surface temperatures rose well above normal. Some reefs are recovering, but scientists say that between 50% and 70% of the world's coral reefs are now endangered and predict that global warming could exacerbate this trend.


Open ocean

The open ocean is relatively unproductive because of a lack of nutrients, yet because it is so vast, in total it produces the most primary productivity. The open ocean is separated into different zones, and the different zones each have different ecologies. Zones which vary according to their depth include the epipelagic, mesopelagic,
bathypelagic The bathypelagic zone or bathyal zone (from Greek βαθύς (bathýs), deep) is the part of the open ocean that extends from a depth of below the ocean surface. It lies between the mesopelagic above, and the abyssopelagic below. The bathypelagi ...
, abyssopelagic, and hadopelagic zones. Zones which vary by the amount of light they receive include the
photic The photic zone, euphotic zone, epipelagic zone, or sunlight zone is the uppermost layer of a body of water that receives sunlight, allowing phytoplankton to perform photosynthesis. It undergoes a series of physical, chemical, and biological pro ...
and aphotic zones. Much of the aphotic zone's energy is supplied by the open ocean in the form of detritus.


Deep sea and trenches

The deepest recorded oceanic trench measured to date is the Mariana Trench, near the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
, in the
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the conti ...
at . At such depths, water pressure is extreme and there is no sunlight, but some life still exists. A white flatfish, a shrimp and a jellyfish were seen by the American crew of the bathyscaphe ''
Trieste Trieste ( , ; sl, Trst ; german: Triest ) is a city and seaport in northeastern Italy. It is the capital city, and largest city, of the autonomous region of Friuli Venezia Giulia, one of two autonomous regions which are not subdivided into pr ...
'' when it dove to the bottom in 1960. In general, the deep sea is considered to start at the aphotic zone, the point where sunlight loses its power of transference through the water. Many life forms that live at these depths have the ability to create their own light known as bio-luminescence. Marine life also flourishes around seamounts that rise from the depths, where fish and other sea life congregate to spawn and feed. Hydrothermal vents along the
mid-ocean ridge A mid-ocean ridge (MOR) is a seafloor mountain system formed by plate tectonics. It typically has a depth of about and rises about above the deepest portion of an ocean basin. This feature is where seafloor spreading takes place along a div ...
spreading centers act as oases, as do their opposites, cold seeps. Such places support unique
biome A biome () is a biogeographical unit consisting of a biological community that has formed in response to the physical environment in which they are found and a shared regional climate. Biomes may span more than one continent. Biome is a broader ...
s and many new microbes and other lifeforms have been discovered at these locations.


Marine life

In biology many phyla, families and genera have some species that live in the sea and others that live on land. Marine biology classifies species based on the environment rather than on taxonomy. For this reason marine biology encompasses not only organisms that live only in a marine environment, but also other organisms whose lives revolve around the sea.


Microscopic life

As inhabitants of the largest environment on Earth, microbial marine systems drive changes in every global system. Microbes are responsible for virtually all the photosynthesis that occurs in the ocean, as well as the cycling of
carbon Carbon () is a chemical element with the symbol C and atomic number 6. It is nonmetallic and tetravalent—its atom making four electrons available to form covalent chemical bonds. It belongs to group 14 of the periodic table. Carbon ma ...
, nitrogen,
phosphorus Phosphorus is a chemical element with the symbol P and atomic number 15. Elemental phosphorus exists in two major forms, white phosphorus and red phosphorus, but because it is highly reactive, phosphorus is never found as a free element on Ea ...
and other
nutrients A nutrient is a substance used by an organism to survive, grow, and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi, and protists. Nutrients can be incorporated into cells for metabolic purposes or excr ...
and trace elements. Microscopic life undersea is incredibly diverse and still poorly understood. For example, the role of
virus A virus is a submicroscopic infectious agent that replicates only inside the living cells of an organism. Viruses infect all life forms, from animals and plants to microorganisms, including bacteria and archaea. Since Dmitri Ivanovsk ...
es in marine ecosystems is barely being explored even in the beginning of the 21st century. The role of phytoplankton is better understood due to their critical position as the most numerous primary producers on Earth. Phytoplankton are categorized into cyanobacteria (also called blue-green algae/bacteria), various types of algae (red, green, brown, and yellow-green), diatoms, dinoflagellates,
euglenoids Euglenids (euglenoids, or euglenophytes, formally Euglenida/Euglenoida, ICZN, or Euglenophyceae, ICBN) are one of the best-known groups of flagellates, which are excavate eukaryotes of the phylum Euglenophyta and their cell structure is typical ...
,
coccolithophorid Coccolithophores, or coccolithophorids, are single celled organisms which are part of the phytoplankton, the autotrophic (self-feeding) component of the plankton community. They form a group of about 200 species, and belong either to the kingd ...
s, cryptomonads, chrysophytes,
chlorophyte Chlorophyta or Prasinophyta is a taxon of green algae informally called chlorophytes. The name is used in two very different senses, so care is needed to determine the use by a particular author. In older classification systems, it refers to a ...
s,
prasinophyte The prasinophytes are a group of unicellular green algae. Prasinophytes mainly include marine planktonic species, as well as some freshwater representatives.Sym, S. D. and Pienaar, R. N. 1993. The class Prasinophyceae. In Round, F. E. and Chapm ...
s, and silicoflagellates. Zooplankton tend to be somewhat larger, and not all are microscopic. Many Protozoa are zooplankton, including dinoflagellates, zooflagellates, foraminiferans, and radiolarians. Some of these (such as dinoflagellates) are also phytoplankton; the distinction between plants and animals often breaks down in very small organisms. Other zooplankton include cnidarians, ctenophores,
chaetognaths The Chaetognatha or chaetognaths (meaning ''bristle-jaws'') are a phylum of predatory marine worms that are a major component of plankton worldwide. Commonly known as arrow worms, about 20% of the known Chaetognatha species are benthic, and can ...
, molluscs,
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chiti ...
s, urochordates, and annelids such as polychaetes. Many larger animals begin their life as zooplankton before they become large enough to take their familiar forms. Two examples are fish larvae and sea stars (also called starfish).


Plants and algae

Microscopic algae and plants provide important habitats for life, sometimes acting as hiding places for larval forms of larger fish and foraging places for invertebrates. Algal life is widespread and very diverse under the ocean. Microscopic photosynthetic algae contribute a larger proportion of the world's photosynthetic output than all the terrestrial forests combined. Most of the
niche Niche may refer to: Science *Developmental niche, a concept for understanding the cultural context of child development * Ecological niche, a term describing the relational position of an organism's species *Niche differentiation, in ecology, the ...
occupied by sub plants on land is actually occupied by macroscopic algae in the ocean, such as '' Sargassum'' and kelp, which are commonly known as seaweeds that create kelp forests. Plants that survive in the sea are often found in shallow waters, such as the seagrasses (examples of which are eelgrass, '' Zostera'', and turtle grass, ''Thalassia''). These plants have adapted to the high salinity of the ocean environment. The intertidal zone is also a good place to find plant life in the sea, where mangroves or cordgrass or
beach grass ''Ammophila'' (synonymous with ''Psamma'' P. Beauv.) is a genus of flowering plants consisting of two or three very similar species of grasses. The common names for these grasses include marram grass, bent grass, and beachgrass. These grasses ar ...
might grow.


Invertebrates

As on land, invertebrates make up a huge portion of all life in the sea. Invertebrate sea life includes
Cnidaria Cnidaria () is a phylum under kingdom Animalia containing over 11,000 species of aquatic animals found both in freshwater and marine environments, predominantly the latter. Their distinguishing feature is cnidocytes, specialized cells that ...
such as
jellyfish Jellyfish and sea jellies are the informal common names given to the medusa-phase of certain gelatinous members of the subphylum Medusozoa, a major part of the phylum Cnidaria. Jellyfish are mainly free-swimming marine animals with umbre ...
and sea anemones; Ctenophora;
sea worm Sea worm may refer to one or several of the following phyla: See also * Marine worm Any worm that lives in a marine environment is considered a marine worm. Marine worms are found in several different phyla, including the Platyhelminthes, ...
s including the
phyla Phyla, the plural of ''phylum'', may refer to: * Phylum, a biological taxon between Kingdom and Class * by analogy, in linguistics, a large division of possibly related languages, or a major language family which is not subordinate to another Phy ...
Platyhelminthes, Nemertea, Annelida, Sipuncula, Echiura,
Chaetognatha The Chaetognatha or chaetognaths (meaning ''bristle-jaws'') are a phylum of predatory marine worms that are a major component of plankton worldwide. Commonly known as arrow worms, about 20% of the known Chaetognatha species are benthic, and ca ...
, and Phoronida; Mollusca including shellfish, squid, octopus; Arthropoda including Chelicerata and Crustacea; Porifera; Bryozoa;
Echinodermata An echinoderm () is any member of the phylum Echinodermata (). The adults are recognisable by their (usually five-point) radial symmetry, and include starfish, brittle stars, sea urchins, sand dollars, and sea cucumbers, as well as the s ...
including starfish; and
Urochordata A tunicate is a marine invertebrate animal, a member of the subphylum Tunicata (). It is part of the Chordata, a phylum which includes all animals with dorsal nerve cords and notochords (including vertebrates). The subphylum was at one time cal ...
including sea squirts or
tunicate A tunicate is a marine invertebrate animal, a member of the subphylum Tunicata (). It is part of the Chordata, a phylum which includes all animals with dorsal nerve cords and notochords (including vertebrates). The subphylum was at one t ...
s. Invertebrates have no backbone. There are over a million species.


Fungi

Over 10,000 species of
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately fr ...
are known from marine environments. These are parasitic on marine algae or animals, or are saprobes on algae, corals, protozoan cysts, sea grasses, wood and other substrata, and can also be found in sea foam. Spores of many species have special appendages which facilitate attachment to the substratum. A very diverse range of unusual secondary metabolites is produced by marine fungi.


Vertebrates


Fish

A reported 33,400 species of fish, including bony and cartilaginous fish, had been described by 2016, more than all other vertebrates combined. About 60% of fish species live in saltwater.


Reptiles

Reptiles which inhabit or frequent the sea include sea turtles,
sea snake Sea snakes, or coral reef snakes, are elapid snakes that inhabit marine environments for most or all of their lives. They belong to two subfamilies, Hydrophiinae and Laticaudinae. Hydrophiinae also includes Australasian terrestrial snakes, w ...
s, terrapins, the marine iguana, and the
saltwater crocodile The saltwater crocodile (''Crocodylus porosus'') is a crocodilian native to saltwater habitats and brackish wetlands from India's east coast across Southeast Asia and the Sundaic region to northern Australia and Micronesia. It has been l ...
. Most extant marine reptiles, except for some sea snakes, are oviparous and need to return to land to lay their eggs. Thus most species, excepting sea turtles, spend most of their lives on or near land rather than in the ocean. Despite their marine adaptations, most sea snakes prefer shallow waters nearby land, around islands, especially waters that are somewhat sheltered, as well as near estuaries.Stidworthy J. 1974. Snakes of the World. Grosset & Dunlap Inc. 160 pp. . tp://ftp.fao.org/docrep/fao/009/y0870e/y0870e65.pdf Sea snakesa
Food and Agriculture Organization of the United Nations
Accessed 7 August 2007.
Some extinct marine reptiles, such as ichthyosaurs, evolved to be viviparous and had no requirement to return to land.


Birds

Birds adapted to living in the marine environment are often called seabirds. Examples include albatross, penguins, gannets, and
auk An auk or alcid is a bird of the family Alcidae in the order Charadriiformes. The alcid family includes the murres, guillemots, auklets, puffins, and murrelets. The word "auk" is derived from Icelandic ''álka'', from Old Norse ''alka'' (a ...
s. Although they spend most of their lives in the ocean, species such as gulls can often be found thousands of miles inland.


Mammals

There are five main types of marine mammals, namely cetaceans (
toothed whale The toothed whales (also called odontocetes, systematic name Odontoceti) are a parvorder of cetaceans that includes dolphins, porpoises, and all other whales possessing teeth, such as the beaked whales and sperm whales. Seventy-three species of t ...
s and baleen whales); sirenians such as manatees;
pinniped Pinnipeds (pronounced ), commonly known as seals, are a widely distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammals. They comprise the extant families Odobenidae (whose only living member is the ...
s including seals and the walrus; sea otters; and the polar bear. All are air-breathing, and while some such as the sperm whale can dive for prolonged periods, all must return to the surface to breathe.


Subfields

The marine ecosystem is large, and thus there are many sub-fields of marine biology. Most involve studying specializations of particular animal groups, such as phycology,
invertebrate zoology Invertebrate zoology is the subdiscipline of zoology that consists of the study of invertebrates, animals without a backbone (a structure which is found only in fish, amphibians, reptiles, birds and mammals). Invertebrates are a vast and very ...
and ichthyology. Other subfields study the physical effects of continual immersion in sea water and the ocean in general, adaptation to a salty environment, and the effects of changing various oceanic properties on marine life. A subfield of marine biology studies the relationships between oceans and ocean life, and global warming and environmental issues (such as
carbon dioxide Carbon dioxide ( chemical formula ) is a chemical compound made up of molecules that each have one carbon atom covalently double bonded to two oxygen atoms. It is found in the gas state at room temperature. In the air, carbon dioxide is t ...
displacement). Recent marine
biotechnology Biotechnology is the integration of natural sciences and engineering sciences in order to achieve the application of organisms, cells, parts thereof and molecular analogues for products and services. The term ''biotechnology'' was first used ...
has focused largely on marine biomolecules, especially
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, res ...
s, that may have uses in medicine or engineering. Marine environments are the home to many exotic biological materials that may inspire
biomimetic material Biomimetic materials are materials developed using inspiration from nature. This may be useful in the design of composite materials. Natural structures have inspired and innovated human creations. Notable examples of these natural structures includ ...
s.


Related fields

Marine biology is a branch of biology. It is closely linked to oceanography, especially biological oceanography, and may be regarded as a sub-field of marine science. It also encompasses many ideas from
ecology Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overl ...
.
Fisheries science Fisheries science is the academic discipline of managing and understanding fisheries. It is a multidisciplinary science, which draws on the disciplines of limnology, oceanography, freshwater biology, marine biology, meteorology, conservation, ...
and
marine conservation Marine conservation, also known as ocean conservation, is the protection and preservation of ecosystems in oceans and seas through planned management in order to prevent the over-exploitation of these marine resources. Marine conservation is i ...
can be considered partial offshoots of marine biology (as well as environmental studies). Marine Chemistry, Physical oceanography and Atmospheric sciences are closely related to this field.


Distribution factors

An active research topic in marine biology is to discover and map the life cycles of various species and where they spend their time. Technologies that aid in this discovery include pop-up satellite archival tags, acoustic tags, and a variety of other data loggers. Marine biologists study how the
ocean current An ocean current is a continuous, directed movement of sea water generated by a number of forces acting upon the water, including wind, the Coriolis effect, breaking waves, cabbeling, and temperature and salinity differences. Depth conto ...
s, tides and many other oceanic factors affect ocean life forms, including their growth, distribution and well-being. This has only recently become technically feasible with advances in GPS and newer underwater visual devices. Most ocean life breeds in specific places, nests or not in others, spends time as juveniles in still others, and in maturity in yet others. Scientists know little about where many species spend different parts of their life cycles especially in the infant and juvenile years. For example, it is still largely unknown where juvenile sea turtles and some year-1
shark Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachi ...
s travel. Recent advances in underwater tracking devices are illuminating what we know about marine organisms that live at great Ocean depths. The information that pop-up satellite archival tags give aids in certain time of the year fishing closures and development of a
marine protected area Marine protected areas (MPA) are protected areas of seas, oceans, estuaries or in the US, the Great Lakes. These marine areas can come in many forms ranging from wildlife refuges to research facilities. MPAs restrict human activity for a con ...
. This data is important to both scientists and fishermen because they are discovering that by restricting commercial fishing in one small area they can have a large impact in maintaining a healthy fish population in a much larger area.


History

The study of marine biology dates back to
Aristotle Aristotle (; grc-gre, Ἀριστοτέλης ''Aristotélēs'', ; 384–322 BC) was a Greek philosopher and polymath during the Classical period in Ancient Greece. Taught by Plato, he was the founder of the Peripatetic school of ...
(384–322 BC), who made many observations of life in the sea around Lesbos, laying the foundation for many future discoveries. In 1768,
Samuel Gottlieb Gmelin Samuel George Gottlieb Gmelin (4 July 1744 – 27 July 1774) was a German physician, botanist, and explorer. Background Gmelin was born at Tübingen as part of a well-known family of naturalists. His father was Johann Conrad Gmelin, an apotheca ...
(1744–1774) published the ''Historia Fucorum'', the first work dedicated to marine algae and the first book on marine biology to use the new
binomial nomenclature In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name compos ...
of Linnaeus. It included elaborate illustrations of seaweed and marine algae on folded leaves. The British naturalist
Edward Forbes Edward Forbes FRS, FGS (12 February 1815 – 18 November 1854) was a Manx naturalist. In 1846, he proposed that the distributions of montane plants and animals had been compressed downslope, and some oceanic islands connected to the mainlan ...
(1815–1854) is generally regarded as the founder of the science of marine biology. The pace of oceanographic and marine biology studies quickly accelerated during the course of the 19th century. The observations made in the first studies of marine biology fueled the age of discovery and exploration that followed. During this time, a vast amount of knowledge was gained about the life that exists in the oceans of the world. Many voyages contributed significantly to this pool of knowledge. Among the most significant were the voyages of where
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended ...
came up with his theories of evolution and on the formation of
coral reefs A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. ...
. Another important expedition was undertaken by HMS ''Challenger'', where findings were made of unexpectedly high species diversity among
fauna Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is '' flora'', and for fungi, it is '' funga''. Flora, fauna, funga and other forms of life are collectively referred to as '' biota''. ...
stimulating much theorizing by population ecologists on how such varieties of life could be maintained in what was thought to be such a hostile environment. This era was important for the history of marine biology but naturalists were still limited in their studies because they lacked technology that would allow them to adequately examine species that lived in deep parts of the oceans. The creation of marine laboratories was important because it allowed marine biologists to conduct research and process their specimens from expeditions. The oldest marine laboratory in the world, Station biologique de Roscoff, was established in Concarneau, France founded by the College of France in 1859. In the United States, the Scripps Institution of Oceanography dates back to 1903, while the prominent
Woods Hole Oceanographic Institute The Woods Hole Oceanographic Institution (WHOI, acronym pronounced ) is a private, nonprofit research and higher education facility dedicated to the study of marine science and engineering. Established in 1930 in Woods Hole, Massachusetts, it ...
was founded in 1930. The development of technology such as sound navigation ranging, scuba diving gear, submersibles and remotely operated vehicles allowed marine biologists to discover and explore life in deep oceans that was once thought to not exist.


See also

*
Acoustic ecology Acoustic ecology, sometimes called ecoacoustics or soundscape studies, is a discipline studying the relationship, mediated through sound, between human beings and their environment. Acoustic ecology studies started in the late 1960s with R. Mur ...
* Aquaculture *
Bathymetry Bathymetry (; ) is the study of underwater depth of ocean floors ('' seabed topography''), lake floors, or river floors. In other words, bathymetry is the underwater equivalent to hypsometry or topography. The first recorded evidence of water ...
* Biological oceanography * Effects of climate change on oceans * Freshwater biology * Modular ocean model * Oceanic basin * Oceanic climate * Phycology


Lists

* Glossary of ecology * Index of biology articles * Large marine ecosystem *
List of ecologists This is a list of notable ecologists. A-D * John Aber (USA) * Aziz Ab'Saber ( Brazil) * Charles Christopher Adams (USA) * Warder Clyde Allee (USA) * Herbert G. Andrewartha ( Australia) * Sarah Martha Baker ( UK) * Fakhri A. Bazzaz (USA) * ...
* List of marine biologists *
List of marine ecoregions (WWF) The following is a list of marine ecoregions, as defined by the WWF and The Nature Conservancy The WWF/Nature Conservancy scheme groups the individual ecoregions into 12 marine realms, which represent the broad latitudinal divisions of polar, tem ...
* Outline of biology *
Outline of ecology The following outline is provided as an overview of and topical guide to ecology: Ecology – scientific study of the distribution and abundance of living organisms and how the distribution and abundance are affected by interactions bet ...


References


Further references

* Morrissey J and Sumich J (2011
''Introduction to the Biology of Marine Life''
Jones & Bartlett Publishers. . * Mladenov, Philip V., Marine Biology: A Very Short Introduction, 2nd edn (Oxford, 2020; online edn, Very Short Introductions online, Feb. 2020), http://dx.doi.org/10.1093/actrade/9780198841715.001.0001, accessed 21 Jun. 2020.


External links


Smithsonian Ocean Portal

Marine Conservation Society
*
Marine Ecology - an evolutionary perspective

Free special issue: Marine Biology in Time and Space

Creatures of the deep ocean
– ''National Geographic'' documentary, 2010.
Exploris


- From the University of Washington Library
Marine Training Portal
- Portal grouping training initiatives in the field of Marine Biology {{DEFAULTSORT:Marine Biology Biological oceanography Fisheries science Oceanographical terminology