Tunica (biology)

In biology, a tunica (, ; ) is a layer, coat, sheath, or similar covering. The word came to English from the New Latin of science and medicine. Its literal sense is about the same as that of the word ''tunic'', with which it is cognate. In biology one of its senses used to be the taxonomic name of a genus of plants, but the nomenclature has been revised and those plants are now included in the genus ''Petrorhagia''.

In modern biology in general, ''tunica'' occurs as a technical or anatomical term mainly in botany and zoology. It usually refers to membranous structures that line or cover particular organs. In many such contexts ''tunica'' is used interchangeably with ''tunic'' according to preference. An organ or organism that has a tunic(a) may be said to be ''tunicate'', as in a ''tunicate bulb''. This adjective ''tunicate'' is not to be confused with the noun ''tunicate'', which refers to a member of the subphylum '' Tunicata''.

Botanical and related usages

In botany there are several contexts for the term.
*As a general, but not comprehensive, descriptive term in botanical anatomy John Lindley in the 19th century defined tunica as "any loose membranous skin not formed from epidermis".Jackson, Benjamin, Daydon; A Glossary of Botanic Terms with their Derivation and Accent; Published by Gerald Duckworth & Co. London, 4th ed 1928
*The apical meristem, in particular in Angiosperms, has an outer layer of cells called the tunica; its role in growth and development differs from that of the inner meristem, or ''corpus''.
*Bulbs and corms often have protective outer coverings of dead material. Particularly in fields such as horticulture and taxonomic description, such bulbs and corms are said to be "tunicate" — having tunics. Usually such a tunic is derived from the bases of sacrificial dead leaves, often cataphylls specialised for the purpose and dying in place. Such tunics may be delicate, brittle membranes such as those around the bulbs of onions, but the many variations reflect a wide range of adaptations in various species. Many of the larger Amaryllidaceae in particular, for example '' Boophone'' species, accumulate thick layers of rot-resistant leaf-base material around their bulbs. Again, corms of Iridaceous species, such as some Watsonias and the larger species of Gladiolus, accumulate thick, reticulated, fibrous or woody defences.
*More generally than in describing tunics of bulbs etc., any leaf-sheath or protective bract remaining attached to the plant after the leaf has died may be called a tunica or tunic.
*The testa or spermoderm of a seed is sometimes called the tunic, especially in older books.
*In fungi the peridium may be referred to as the tunica.

Zoological usages

As an anatomical or morphological reference in zoology, ''tunica'' has a range of applications to membranous structures in anatomy, including human anatomy. Such structures are generally coverings or capsules of organs, but also may be linings of cavities. In some cases, such as the walls of macroscopic blood vessels, layers of the tissue of the walls, whether inner, intermediate, or outer, are called ''tunica'' of one kind or another. Examples follow, but neither the list nor the discussions are exhaustive.
*In arteries and veins the inner wall is the ''tunica intima'', the outer wall is the ''tunica adventitia'', and they are separated by the ''tunica media''. More generally, the ''tunica adventitia'' or simply the "adventitia" is the outermost connective tissue covering round any internal organ.Junqueira, L. Carlos; Carneiro, Jose; Kelly, Robert O. Basic Histology, Publisher: Prentice Hall 1995
*'' Tunica albuginea'' is a general term for a tunic of whitish connective tissue. In human anatomy it generally is applied in three contexts: the fibrous sheath of the ''corpora cavernosa penis'', the fibrous tunic of the testis, and the connective tissue tunic around the ovaries.
*'' Tunica dartos'' is a muscular tunic beneath the skin of the scrotum. It is one of the mechanisms for retraction of the testes, largely for temperature control.Amenta, Peter S. Histology: From Normal Microanatomy to Pathology. Publisher: Piccin Nuova Libraria 1997
*'' Tunica fibrosa oculi'', is the fibrous tunic of the eyeball, the outer layer that includes both the cornea and sclera.
*'' Tunica mucosa'' is the predominant Latin rendering of "mucous membrane" in English-speaking countries, though ''membrana mucosa'' also is acceptable and in some European languages is preferred. The name may be qualified in referring to particular regions of the body, such as '' tunica mucosa oris'' for the oral mucosa, and '' tunica mucosa uteri'' for the endometrium.
*'' Tunica muscularis'' refers to muscular layers in the walls of hollow organs such as the gut, where they are required for peristalsis, and sometimes for sphincters.
*'' Tunica serosa'' is the serous membrane
*'' Tunica vaginalis'' is the serous covering of the testis within the scrotum.
*''Tunica vasculosa'' could refer to any tunic rich in blood vessels. In human anatomy it generally refers either to ''Tunica vasculosa lentis'' or to '' Tunica vasculosa testis''Shu-Xin Zhang. An Atlas of Histology. Publisher: Springer 1999

References

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Animal anatomy
Plant anatomy