Oscines (augury)
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A songbird is a
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
belonging to the suborder Passeri of the perching birds (
Passeriformes A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by th ...
). Another name that is sometimes seen as the scientific or vernacular name is Oscines, from
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
''oscen'', "songbird". The Passeriformes contains 5000 or so speciesEdwards, Scott V. and John Harshman. 2013. Passeriformes. Perching Birds, Passerine Birds. Version 06 February 2013 (under construction). http://tolweb.org/Passeriformes/15868/2013.02.06 in The Tree of Life Web Project, http://tolweb.org/ ccessed 2017/12/11 found all over the world, in which the vocal organ typically is developed in such a way as to produce a diverse and elaborate bird song. Songbirds form one of the two major lineages of extant perching birds (~4000 species), the other being the
Tyranni The Tyranni (suboscines) are a suborder of passerine birds that includes more than 1,000 species, the large majority of which are South American. It is named after the type genus '' Tyrannus''. These have a different anatomy of the syrinx mus ...
(~1000 species), which are most diverse in the
Neotropics The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...
and absent from many parts of the world. The Tyranni have a simpler syrinx musculature, and while their vocalizations are often just as complex and striking as those of songbirds, they are altogether more mechanical sounding. There is a third perching bird lineage, the
Acanthisitti The New Zealand wrens are a family (Acanthisittidae) of tiny passerines endemic to New Zealand. They were represented by seven Holocene species in four or five genera, although only two species in two genera survive today. They are understood to ...
from
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
, of which only two species remain alive today. Some evidence suggests that songbirds evolved 50 million years ago in the part of Gondwana that later became
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
, Sri Lanka, Australia, New Zealand,
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; id, Papua, or , historically ) is the world's second-largest island with an area of . Located in Oceania in the southwestern Pacific Ocean, the island is separated from Australia by the wide Torr ...
and
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest cont ...
, before spreading around the world.


Description

The song in this clade is essentially territorial, because it communicates the identity and whereabouts of an individual to other birds, and also signals sexual intentions. Sexual selection among songbirds is highly based on mimetic vocalization. Female preference has shown in some populations to be based on the extent of a male's song repertoire. The larger a male's repertoire, the more females a male individual attracts. It is not to be confused with bird calls that are used for alarms and contact and are especially important in birds that feed or migrate in flocks. While almost all living birds give calls of some sort, well-developed songs are only given by a few lineages outside the songbirds. And still, not all songbirds proffer a call that is distinctly melodious. Songbirds do, however, possess a highly developed vocal organ, the syrinx, that enables their sonorous activity. This organ, also known as a song box, can be found where the windpipe meets diverging bronchial tubes which lead to the lungs. The organ is a solid, bony structure lined with a film of membranes which air passes through as the songbird calls. While the song boxes of songbirds vary in size and intricacy, this does not necessarily determine the songbird's ability to voice their song. Researchers believe this has more to do with the length of the windpipe. Other birds (especially non-passeriforms) sometimes have
songs A song is a musical composition intended to be performed by the human voice. This is often done at distinct and fixed pitches (melodies) using patterns of sound and silence. Songs contain various forms, such as those including the repetition ...
to attract mates or hold territory, but these are usually simple and repetitive, lacking the variety of many oscine songs. The monotonous repetition of the
common cuckoo The common cuckoo (''Cuculus canorus'') is a member of the cuckoo order of birds, Cuculiformes, which includes the roadrunners, the anis and the coucals. This species is a widespread summer migrant to Europe and Asia, and winters in Africa. I ...
or
little crake The little crake (''Zapornia parva'') is a very small waterbird of the family Rallidae. ''parva'' is Latin for "small". Its breeding habitat is reed beds in Europe, mainly in the east, and just into western Asia. This species is migratory, wi ...
can be contrasted with the variety of a
nightingale The common nightingale, rufous nightingale or simply nightingale (''Luscinia megarhynchos''), is a small passerine bird best known for its powerful and beautiful song. It was formerly classed as a member of the thrush family Turdidae, but is no ...
or marsh warbler. On the other hand, although many songbirds have songs that are pleasant to the human ear, this is not invariably the case. Many members of the
crow A crow is a bird of the genus '' Corvus'', or more broadly a synonym for all of ''Corvus''. Crows are generally black in colour. The word "crow" is used as part of the common name of many species. The related term "raven" is not pinned scientifica ...
family (
Corvidae Corvidae is a cosmopolitan family of oscine passerine birds that contains the crows, ravens, rooks, jackdaws, jays, magpies, treepies, choughs, and nutcrackers. In colloquial English, they are known as the crow family or corvids. Cu ...
) communicate with croaks or screeches, which sound harsh to humans. Even these, however, have a song of sorts, a softer twitter that is given between courting partners. And even though some
parrot Parrots, also known as psittacines (), are birds of the roughly 398 species in 92 genera comprising the order Psittaciformes (), found mostly in tropical and subtropical regions. The order is subdivided into three superfamilies: the Psittacoide ...
s (which are not songbirds) can be taught to repeat human speech, vocal mimicry among birds is almost completely restricted to songbirds, some of which (such as the
lyrebird A lyrebird is either of two species of ground-dwelling Australian birds that compose the genus ''Menura'', and the family Menuridae. They are most notable for their impressive ability to mimic natural and artificial sounds from their environ ...
s or the aptly-named
mockingbird Mockingbirds are a group of New World passerine birds from the family Mimidae. They are best known for the habit of some species mimicking the songs of other birds and the sounds of insects and amphibians, often loudly and in rapid succession. ...
s) excel in imitating the sounds of other birds or even environmental noises. The birds from higher altitudes have evolved thicker downs (also known as jackets) to protect themselves from the changes in climate. Their feathers have outer and inner portions, with the lower down being fluffier and warmer to provide increased warmth.


Song repertoire and courtship

Sexual selection can be broken down into several different studies regarding different aspects of a bird's song. As a result, song can vary even within a single species. Many believe that song repertoire and cognition have a direct relationship. However, a study published in 2013 has shown that all cognitive ability may not be directly related to the song repertoire of a songbird. Specifically, spatial learning is said to have an inverse relationship with song repertoire. So for example, this would be an individual who does not migrate as far as others in the species, but has a better song repertoire. This suggests an evolutionary trade-off between possible alleles. With natural selection choosing traits best fit for reproductive success there could be a trade off in either direction depending on which trait would produce a higher fitness at that time period. Song repertoire can be attributed to male songbirds as it is one of the main mechanisms of courtship. Song repertoires differ from male individual to male individual and species to species. Some species may typically have large repertoires while others may have significantly smaller ones. Mate choice in female songbirds is a significant realm of study as song abilities are continuously evolving. Males often sing to assert their dominance over other males in competition for a female, sometimes in lieu of a combative episode, and to arouse the female by announcing a readiness to mate. Though less frequent, females have also been known to sing and occasionally in duet with a mate as an affirmation of their partnership. While some will sing their song from a familiar perch, other species common to grasslands will sing a familiar song each time they fly. Currently there have been numerous studies involving songbird repertoires, unfortunately, there has yet been concrete evidence to confirm that every songbird species prefers larger repertoires. A conclusion can be made that it can vary between species on whether a larger repertoire is connected to better fitness. With this conclusion, it can be inferred that evolution via natural selection, or sexual selection, favors the ability to retain larger repertoires for these certain species as it leads to higher reproductive success. During times of courtship, it is said that male songbirds increase their repertoire by mimicking other species songs. The better the mimicking ability, retaining ability, and the quantity of other species mimicked has been proven to have a positive relationship with mating success. Female preferences cause the constant improvement of accuracy and presentation of the copied songs. Another theory known as the "song-sharing hypothesis" suggests that females prefer simpler, more homogenous songs that signal a male of familiar territory. As birdsong can be broken into regional dialects through this process of mimicry, the foreign song of a newcomer suggests the lack of territorial possession. This can be costly in the wake of territorial conflicts between disparate songbird populations and may compel a female to prefer a male spouting a familiar song of the area.


Taxonomy and systematics

Sibley and Alquist divided songbirds into two "
parvorder Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and ...
s",
Corvida The "Corvida" were one of two "parvorders" contained within the suborder Passeri, as proposed in the Sibley-Ahlquist taxonomy, the other being Passerida. Standard taxonomic practice would place them at the rank of infraorder. More recent rese ...
and
Passerida Passerida is, under the Sibley-Ahlquist taxonomy, one of two parvorders contained within the suborder Passeri (standard taxonomic practice would place them at the rank of infraorder). While more recent research suggests that its sister parvorde ...
(standard taxonomic practice would rank these as
infraorder Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and ...
s), distributed in Australo-Papua and
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago ...
respectively.Selvatti, A.P. ''et al''. (2015
A Paleogene origin for crown passerines and the diversification of the Oscines in the New World
''Molecular Phylogenetics and Evolution'', 88:1-15.
Subsequent molecular studies, however, show this treatment to be somewhat erroneous. Passerida is a highly diverse lineage, uniting over one third of all bird species to include (in 2015) 3885 species). These are divided into three major superfamilies (though not exactly corresponding to the Sibley-Ahlquist arrangement), in addition to some minor lineages. In contrast, Sibley & Alquist's "Corvida" is a phylogenetic
grade Grade most commonly refers to: * Grade (education), a measurement of a student's performance * Grade, the number of the year a student has reached in a given educational stage * Grade (slope), the steepness of a slope Grade or grading may also ref ...
, and an artefact of the
phenetic In biology, phenetics ( el, phainein – to appear) , also known as taximetrics, is an attempt to classify organisms based on overall similarity, usually in morphology or other observable traits, regardless of their phylogeny or evolutionary re ...
methodology. The bulk of the "Corvida" make up the large clade Corvides (812 species as of 2015), which is a sister group to the Passerida. The remaining 15 oscine families (343 species in 2015) form a series of basally branching sister groups to the Corvoid - Passerid clade. All of these groups, which form at least six successively branching basal clades, are found exclusively or predominantly in Australasia. Australian endemics are also prominent among basal lineages in both Corvoids and Passerids, suggesting that songbirds originated and diverged in Australia. Scrub-birds and lyrebirds, of which there are just two species of each, represent the oldest lineage of songbirds on Earth. The rufous scrubbird, ''Atrichornis rufescens'', is essentially confined to the Gondwana Rainforests of Australia World Heritage Area, occurring in both Queensland and New South Wales sections. It is now only found at elevations above 600 m.Chris Cooney, "How songbirds island-hopped their way from Australia to colonise the world", ''The Conversation,'' August 31, 2016.
/ref>


Families

* Menuroidea ** Menuridae:
lyrebird A lyrebird is either of two species of ground-dwelling Australian birds that compose the genus ''Menura'', and the family Menuridae. They are most notable for their impressive ability to mimic natural and artificial sounds from their environ ...
s ** Atrichornithidae: scrub birds * Bowerbirds and Australian treecreepers **
Climacteridae There are seven species of Australasian treecreeper in the passerine bird family Climacteridae. They are medium-small, mostly brown birds with patterning on their underparts, and all are endemic to Australia-New Guinea. They resemble, but are n ...
: Australian
treecreeper The treecreepers are a family, Certhiidae, of small passerine birds, widespread in wooded regions of the Northern Hemisphere and sub-Saharan Africa. The family contains eleven species in two genera, '' Certhia'' and '' Salpornis''. Their plumage ...
s **
Ptilonorhynchidae Bowerbirds () make up the bird family Ptilonorhynchidae. They are renowned for their unique courtship behaviour, where males build a structure and decorate it with sticks and brightly coloured objects in an attempt to attract a mate. The family ...
: bowerbirds * Meliphagoidea: honeyeaters and allies **
Maluridae The Australasian wrens are a family, Maluridae, of small, insectivorous passerine birds endemic to Australia and New Guinea. While commonly known as wrens, they are unrelated to the true wrens. The family comprises 32 species (including sixteen ...
:
fairy-wren The Australasian wrens are a family, Maluridae, of small, insectivorous passerine birds endemic to Australia and New Guinea. While commonly known as wrens, they are unrelated to the true wrens. The family comprises 32 species (including sixteen ...
s,
emu-wren The emu-wrens (''Stipiturus'') are a genus of passerine birds in the Australasian wren family, Maluridae. They are found only in Australia, where they inhabit scrub, heathland and grassland. They are small birds, 12–19 cm long with the ...
s and
grasswren Grasswrens (''Amytornis)'' are a genus of birds in the Australasian wren family, Maluridae. Taxonomy and systematics ''Amytornis'' is the only genus classified within the subfamily Amytornithinae, and form a separate clade than the related fa ...
s **
Meliphagidae The honeyeaters are a large and diverse family, Meliphagidae, of small to medium-sized birds. The family includes the Australian chats, myzomelas, friarbirds, wattlebirds, miners and melidectes. They are most common in Australia and New Gu ...
: true honeyeaters and chats **
Dasyornithidae The bristlebirds are a family of passerine birds, Dasyornithidae. There are three species in one genus, ''Dasyornis''. The family is endemic to the south-east coast and south-west corner of Australia.Del Hoyo, J.; Elliot, A. & Christie D. (edit ...
:
bristlebirds The bristlebirds are a family of passerine birds, Dasyornithidae. There are three species in one genus, ''Dasyornis''. The family is endemic to the south-east coast and south-west corner of Australia.Del Hoyo, J.; Elliot, A. & Christie D. (e ...
** Pardalotidae:
pardalote Pardalotes or peep-wrens are a family, Pardalotidae, of very small, brightly coloured birds native to Australia, with short tails, strong legs, and stubby blunt beaks. This family is composed of four species in one genus, ''Pardalotus'', and seve ...
s **
Acanthizidae The Acanthizidae—known as Australian warblers—are a family of passerine birds which includes gerygones, the thornbills ''Acanthiza'', and the scrubwrens of ''Sericornis''. The Acanthizidae family consists of small to medium passerine birds, ...
: scrubwrens, thornbills, and
gerygone ''Gerygone'' (), the gerygones or peep-warblers, is a genus of bird in the family Acanthizidae. The genus ranges from Southeast Asia through New Guinea and Australia to New Zealand and the Chatham Islands. Most of the species are found in Austr ...
s * Australopapuan babblers ** Pomatostomidae: Australasian babblers * Logrunners ** Orthonychidae: logrunners * Other basal lineages **
Cnemophilidae The satinbirds or cnemophilines, are a family, Cnemophilidae of passerine birds which consists of four species found in the mountain forests of New Guinea. They were originally thought to be part of the birds-of-paradise family Paradisaeidae unti ...
: satinbirds ''
Cnemophilus ''Cnemophilus'' is a genus of satinbirds in the family Cnemophilidae, in which all three species are native to New Guinea mountain slopes and highlands in tropical forests. The generic name ''Cnemophilus'' is Latin for "mountain/slope-lover". ...
'' and '' Loboparadisea'' **
Melanocharitidae The Melanocharitidae, the berrypeckers and longbills, is a small bird family restricted to the forests of New Guinea. The family contains eleven species in four (sometimes three) genera. They are small songbirds with generally dull plumage but a ...
: berrypeckers and longbills ** Callaeidae: New Zealand wattlebirds kokako, saddleback and †
huia The huia ( ; ; ''Heteralocha acutirostris'') is an extinct species of New Zealand wattlebird, endemic to the North Island of New Zealand. The last confirmed sighting of a huia was in 1907, although there was a credible sighting in 1924. It ...
** Notiomystidae: stitchbird * Corvides ** Paramythiidae: tit berrypecker and crested berrypeckers **
Psophodidae Psophodidae is a family of passerine birds native to Australia and nearby areas. It has a complicated taxonomic history and different authors vary in which birds they include in the family. In the strictest sense, it includes only the 5 or 6 spec ...
:
whipbird Psophodidae is a family of passerine birds native to Australia and nearby areas. It has a complicated taxonomic history and different authors vary in which birds they include in the family. In the strictest sense, it includes only the 5 or 6 spec ...
s, jewel-babblers and quail-thrushes ** Platysteiridae:
wattle-eye Platysteiridae is a family of small, stout passerine birds of the African tropics. The family contains the wattle-eyes, batises and shrike-flycatchers. They were previously classed as a subfamily of the Old World flycatchers, Muscicapidae. Thes ...
s and batis ** Malaconotidae: bush-shrikes ** Machaerirynchidae: boatbills ** Vangidae: vangas, woodshrikes, and helmetshrikes ** Pityriasidae:
Bornean bristlehead The Bornean bristlehead (''Pityriasis gymnocephala''), also variously known as the bristled shrike, bald-headed crow or the bald-headed wood-shrike, is the only member of the passerine family (biology), family Pityriasidae and genus ''Pityriasi ...
** Artamidae: butcherbirds,
currawong Currawongs are three species of medium-sized passerine birds belonging to the genus ''Strepera'' in the family Artamidae native to Australia. These are the grey currawong (''Strepera versicolor''), pied currawong (''S. graculina''), and black ...
s and
Australian magpie The Australian magpie (''Gymnorhina tibicen'') is a black and white passerine bird native to Australia and southern New Guinea. Although once considered to be three separate species, it is now considered to be one, with nine recognised su ...
(formerly in Cracticidae) ** Rhagologidae: mottled whistler ** Aegithinidae:
iora : ''For the international organization, see Indian Ocean Rim Association'' : ''For the Australian Aboriginal people of the Sydney region, see Eora'' The ioras are a small family, Aegithinidae, of four passerine bird species found in south and so ...
s ** Campephagidae:
cuckooshrike The cuckooshrikes and allies in the family Campephagidae are small to medium-sized passerine bird species found in the subtropical and tropical Africa, Asia and Australasia. The 93 species are divided into 11 genera. The woodshrikes ('' Tephrodor ...
s and trillers **
Mohouidae ''Mohoua'' is a small genus of three bird species endemic to New Zealand. The scientific name is taken from ''mohua'' – the Māori name for the yellowhead.''"Handbook of Australian, New Zealand and Antarctic Birds"; Volume 7'', edited by Peter ...
: whiteheads ** Neosittidae:
sittella The sittellas are a family, Neosittidae, of small passerine birds found only in Australasia. They resemble nuthatches, but whilst they were considered to be in that family for many years they are now afforded their own family. They do not migrat ...
s ** Eulacestomidae: ploughbill ** Oreoicidae: Australo-Papuan bellbirds ** Pachycephalidae:
whistlers Whistler may refer to: * Someone who whistles Places Canada * Whistler, British Columbia, a resort town ** Whistler railway station ** Whistler Secondary School * Whistler Blackcomb, a ski resort in British Columbia * Whistler Mountain, Briti ...
,
shrike-thrush A shrikethrush, also spelt shrike-thrush, is any one of eleven species of songbird that is a member of the genus ''Colluricincla''. They have nondescript, predominantly brown or grey, plumage, but are accomplished singers, their calls described as ...
es, pitohuis and allies ** Laniidae: shrikes **
Vireonidae The vireos make up a family, Vireonidae, of small to medium-sized passerine birds found in the New World (Canada to Argentina, including Bermuda and the West Indies) and Southeast Asia. "Vireo" is a Latin word referring to a green migratory bir ...
:
vireo The vireos make up a family, Vireonidae, of small to medium-sized passerine birds found in the New World (Canada to Argentina, including Bermuda and the West Indies) and Southeast Asia. "Vireo" is a Latin word referring to a green migrato ...
s ** Oriolidae:
orioles Oriole or Orioles may refer to: Animals * Old World oriole, colorful passerine birds in the family Oriolidae * New World oriole, a group of birds in the family Icteridae Music * The Orioles, an R&B and doo-wop group of the late 1940s and earl ...
, figbirds and † piopio (formerly Turnagridae) ** Dicruridae:
drongo The drongos are a family, Dicruridae, of passerine birds of the Old World tropics. The 30 species in the family are placed in a single genus, ''Dicrurus''. Drongos are mostly black or dark grey, short-legged birds, with an upright stance when ...
s **
Rhipiduridae The family Rhipiduridae are small insectivorous birds of Australasia, Southeast Asia and the Indian subcontinent that includes the fantails and silktails. Taxonomy and systematics There are four genera classified within the family: * Subfamily ...
: fantails ** Monarchidae:
monarch A monarch is a head of stateWebster's II New College DictionarMonarch Houghton Mifflin. Boston. 2001. p. 707. for life or until abdication, and therefore the head of state of a monarchy. A monarch may exercise the highest authority and power i ...
s and allies ** Platylophidae: jayshrike **
Corvidae Corvidae is a cosmopolitan family of oscine passerine birds that contains the crows, ravens, rooks, jackdaws, jays, magpies, treepies, choughs, and nutcrackers. In colloquial English, they are known as the crow family or corvids. Cu ...
:
crow A crow is a bird of the genus '' Corvus'', or more broadly a synonym for all of ''Corvus''. Crows are generally black in colour. The word "crow" is used as part of the common name of many species. The related term "raven" is not pinned scientifica ...
s, magpies, and jays **
Corcoracidae Corcoracidae is a family of passerine birds known as the Australian mudnesters. The family has sometimes been called Struthideidae in the past; however, despite Struthideidae being an older name than Corcoracidae, the latter name takes precedence ...
:
white-winged chough The white-winged chough (''Corcorax melanorhamphos'') is one of only two surviving members of the Australian mud-nest builders family, Corcoracidae, and is the Monotypic taxon, only member of the genus ''Corcorax''. It is native to southern and e ...
and apostlebird ** Melampittidae: melampittas ** Ifritidae: ifritabirds **
Paradisaeidae The birds-of-paradise are members of the family Paradisaeidae of the order Passeriformes. The majority of species are found in eastern Indonesia, Papua New Guinea and eastern Australia. The family has 44 species in 17 genera. The members of this ...
:
birds of paradise The birds-of-paradise are members of the family Paradisaeidae of the order Passeriformes. The majority of species are found in eastern Indonesia, Papua New Guinea and eastern Australia. The family has 44 species in 17 genera. The members of ...
*
Passerida Passerida is, under the Sibley-Ahlquist taxonomy, one of two parvorders contained within the suborder Passeri (standard taxonomic practice would place them at the rank of infraorder). While more recent research suggests that its sister parvorde ...
**
Petroicidae The bird family Petroicidae includes 51 species in 19 genera. All are endemic to Australasia: New Guinea, Australia, New Zealand and numerous Pacific Islands as far east as Samoa. For want of an accurate common name, the family is often called th ...
:
Australasian robin The bird Family (biology), family Petroicidae includes 51 species in 19 genera. All are endemic to Australasia: New Guinea, Australia, New Zealand and numerous Pacific Islands as far east as Samoa. For want of an accurate common name, the family ...
s ** Picathartidae: rockfowl ** Chaetopidae: rockjumpers ** Eupetidae: rail-babbler **
Bombycillidae The waxwings are three species of passerine birds classified in the genus ''Bombycilla''. They are pinkish-brown and pale grey with distinctive smooth plumage in which many body feathers are not individually visible, a black and white eyestri ...
:
waxwing The waxwings are three species of passerine birds classified in the genus ''Bombycilla''. They are pinkish-brown and pale grey with distinctive smooth plumage in which many body feathers are not individually visible, a black and white eyestripe, ...
s and allies **
Ptiliogonatidae The silky-flycatchers are a small family, Ptiliogonatidae, of passerine birds. The family contains only four species in three genera. They were formerly lumped with waxwings and hypocolius in the family Bombycillidae, and they are listed in tha ...
: silky-flycatchers **
Hypocoliidae The grey hypocolius or simply hypocolius (''Hypocolius ampelinus'') is a small passerine bird species. It is the sole member of the genus ''Hypocolius'' and it is placed in a family of its own, the Hypocoliidae. This slender and long tailed bird i ...
: hypocolius **
Dulidae The palmchat (''Dulus dominicus'') is a small, long-tailed passerine bird, the only species in the genus ''Dulus'' and the family Dulidae endemic to the Caribbean island of Hispaniola (split between the Dominican Republic and Haiti). It is rela ...
: palmchat ** †
Mohoidae Mohoidae, also known as the Hawaiian honeyeaters, is a family of Hawaiian species of recently extinct, nectarivorous songbirds in the genera '' Moho'' (ōō) and '' Chaetoptila'' (kioea). These now extinct birds form their own family, represent ...
: some Hawaiian honeyeaters, '' Moho'' and ''
Chaetoptila The kioea (''Chaetoptila angustipluma'') was a endemic Hawaiian bird that became extinct around the mid-19th century. Description The kioea was a large bird, about long, with a long, slightly curved bill. What distinguished the kioea from othe ...
'' genera, not closely related to
Meliphagidae The honeyeaters are a large and diverse family, Meliphagidae, of small to medium-sized birds. The family includes the Australian chats, myzomelas, friarbirds, wattlebirds, miners and melidectes. They are most common in Australia and New Gu ...
**
Hylocitreidae The hylocitrea (''Hylocitrea bonensis''), also known as the yellow-flanked whistler or olive-flanked whistler, is a species of bird that is endemic to montane forests on the Indonesian island of Sulawesi.Boles, W. E. (2007). Yellow-flanked Whistl ...
:
hylocitrea The hylocitrea (''Hylocitrea bonensis''), also known as the yellow-flanked whistler or olive-flanked whistler, is a species of bird that is endemic to montane forests on the Indonesian island of Sulawesi.Boles, W. E. (2007). Yellow-flanked Whistl ...
** Stenostiridae: fairy-flycatcher and allies **
Paridae The tits, chickadees, and titmice constitute the Paridae, a large family of small passerine birds which occur mainly in the Northern Hemisphere and Africa. Most were formerly classified in the genus ''Parus''. Members of this family are common ...
: tits,
chickadee The chickadees are a group of North American birds in the tit family included in the genus '' Poecile''. Species found in North America are referred to as chickadees, while other species in the genus are called tits. They are small-sized bird ...
s, and
titmouse ''Baeolophus'' is a genus of birds in the family Paridae. Its members are commonly known as titmice. All the species are native to North America. In the past, most authorities retained ''Baeolophus'' as a subgenus within the genus ''Parus ...
** Remizidae: penduline-tits **
Nicatoridae The nicators are a genus, ''Nicator'', and family, Nicatoridae, of songbirds endemic to Africa. The genus and family contain three species. Taxonomy The systematic affinities of the genus have been a long-standing mystery. The group was origina ...
:
nicator The nicators are a genus, ''Nicator'', and family, Nicatoridae, of songbirds endemic to Africa. The genus and family contain three species. Taxonomy The systematic affinities of the genus have been a long-standing mystery. The group was origina ...
s ** Panuridae: bearded reedling **
Alaudidae Larks are passerine birds of the family Alaudidae. Larks have a cosmopolitan distribution with the largest number of species occurring in Africa. Only a single species, the horned lark, occurs in North America, and only Horsfield's bush lark occ ...
:
lark Larks are passerine birds of the family Alaudidae. Larks have a cosmopolitan distribution with the largest number of species occurring in Africa. Only a single species, the horned lark, occurs in North America, and only Horsfield's bush lark oc ...
s **
Pycnonotidae The bulbuls are members of a family, Pycnonotidae, of medium-sized passerine songbirds, which also includes greenbuls, brownbuls, leafloves, and bristlebills. The family is distributed across most of Africa and into the Middle East, tropical ...
: bulbuls **
Hirundinidae The swallows, martins, and saw-wings, or Hirundinidae, are a family of passerine songbirds found around the world on all continents, including occasionally in Antarctica. Highly adapted to aerial feeding, they have a distinctive appearance. The ...
:
swallow The swallows, martins, and saw-wings, or Hirundinidae, are a family of passerine songbirds found around the world on all continents, including occasionally in Antarctica. Highly adapted to aerial feeding, they have a distinctive appearance. The ...
s and martins **
Pnoepygidae ''Pnoepyga'' is a genus of passerines endemic to southern and southeastern Asia. Its members are known as cupwings or wren-babblers. The genus contains four species. The genus has long been placed in the babbler family Timaliidae. A 2009 study o ...
: wren-babblers **
Macrosphenidae The African warblers are a newly erected family Macrosphenidae, of songbirds. Most of the species were formerly placed in the Old World warbler family Sylviidae, although one species, the rockrunner, was placed in the babbler family, Timaliidae. ...
: crombecs and African warblers **
Cettiidae Cettiidae is a newly validated family of small insectivorous songbirds ("warblers"), formerly placed in the Old World warbler "wastebin" assemblage. It contains the typical bush warblers (''Cettia'') and their relatives. As a common name, cetti ...
: bush-warblers and allies ** Scotocercidae: streaked scrub-warbler **
Erythrocercidae ''Erythrocercus'' is a genus of birds containing three flycatchers that are found in Africa. The genus is placed in its own family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or af ...
: yellow flycatchers **
Aegithalidae The bushtits or long-tailed tits, are a family, Aegithalidae, of small passerine birds with long tails, compared to their size. The family contains 13 species in three genera, all but one of which are found in Eurasia. Bushtits are active birds, ...
:
long-tailed tit The long-tailed tit (''Aegithalos caudatus''), also named long-tailed bushtit, is a common bird found throughout Europe and the Palearctic. The genus name ''Aegithalos'' was a term used by Aristotle for some European tits, including the long-tail ...
s **
Phylloscopidae Leaf warblers are small insectivorous passerine birds belonging to the genus ''Phylloscopus''. Leaf warblers were formerly included in the Old World warbler family but are now considered to belong to the family Phylloscopidae, introduced in 200 ...
: leaf-warblers and allies. Recently split from Sylviidae. **
Acrocephalidae The Acrocephalidae (the reed warblers, marsh- and tree-warblers, or acrocephalid warblers) are a family of oscine passerine birds, in the superfamily Sylvioidea. The species in this family are usually rather large "warblers". Most are rather p ...
:
reed warbler The ''Acrocephalus'' warblers are small, insectivorous passerine birds belonging to the genus ''Acrocephalus''. Formerly in the paraphyletic Old World warbler assemblage, they are now separated as the namesake of the marsh and tree warbler f ...
s and allies ** Locustellidae: grassbirds and allies ** Donacobiidae:
black-capped donacobius The black-capped donacobius (''Donacobius atricapilla'') is a conspicuous, vocal South American bird. It is found in tropical swamps and wetlands in Argentina, Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Paraguay, Peru, Suriname, ...
**
Bernieridae The Tetrakas and allies are a newly validated family of songbirds. They were formally named Bernieridae in 2010. The family currently consists of eleven species (in eight genera) of small forest birds. These birds are all endemic to Madagascar ...
:
Malagasy warbler The Tetrakas and allies are a newly validated family of songbirds. They were formally named Bernieridae in 2010. The family currently consists of eleven species (in eight genera) of small forest birds. These birds are all endemic to Madagascar. ...
s **
Cisticolidae The family Cisticolidae is a group of about 160 warblers, small passerine birds found mainly in warmer southern regions of the Old World. They were formerly included within the Old World warbler family Sylviidae. This family probably originate ...
:
cisticola __NOTOC__ Cisticolas (pronounced ''sis-TIC-olas'') are a genus of very small insectivorous birds formerly classified in the Old World warbler family Sylviidae, but now usually considered to be in the separate family Cisticolidae, along with other ...
s and allies ** Timaliidae: babblers ** Pellorneidae: ground babblers **
Leiothrichidae The laughingthrushes are a family, Leiothrichidae, of Old World passerine birds. They are diverse in size and coloration. These are birds of tropical areas, with the greatest variety in Southeast Asia and the Indian subcontinent. The entire fam ...
: laughingthrushes and allies **
Sylviidae Sylviidae is a family of passerine birds that includes the typical warblers and a number of babblers formerly placed within the Old World babbler family. They are found in Eurasia and Africa. Taxonomy and systematics The scientific name Sylvii ...
:
Old World warbler Old World warblers are a large group of birds formerly grouped together in the bird family Sylviidae. The family held over 400 species in over 70 genera, and were the source of much taxonomic confusion. Two families were split out initially, the ci ...
s **
Zosteropidae The white-eyes are a family, Zosteropidae, of small passerine birds native to tropical, subtropical and temperate Sub-Saharan Africa, southern and eastern Asia, and Australasia. White-eyes inhabit most tropical islands in the Indian Ocean, the ...
:
white-eye The white-eyes are a family, Zosteropidae, of small passerine birds native to tropical, subtropical and temperate Sub-Saharan Africa, southern and eastern Asia, and Australasia. White-eyes inhabit most tropical islands in the Indian Ocean, the ...
s ** Arcanatoridae: dapple-throat and allies ** Promeropidae: sugarbirds ** Irenidae:
fairy-bluebird The three fairy-bluebirds are small passerine bird species found in forests and plantations in tropical southern Asia and the Philippines. They are the sole members of the genus ''Irena'' and family Irenidae, and are related to the ioras and le ...
s ** Regulidae:
kinglet A kinglet is a small bird in the family Regulidae. Species in this family were formerly classified with the Old World warblers. "Regulidae" is derived from the Latin word ''regulus'' for "petty king" or prince, and refers to the coloured crowns ...
s ** Elachuridae: elachuras **
Hyliotidae The hyliotas are a genus, ''Hyliota'', of passerine bird from Africa. The taxonomic position of the genus has been a longstanding mystery. They have been formerly regarded as Old World warblers in the family Sylviidae, or related to the batises an ...
: hyliotas ** Troglodytidae: wrens ** Polioptilidae:
gnatcatcher The Gnatcatchers are a family of small passerine birds called Polioptilidae. The 21 species occur in North and South America (except for the far south and the high Andean regions). Most species of this mainly tropical and subtropical group are re ...
s **
Sittidae The nuthatches () constitute a genus, ''Sitta'', of small passerine birds belonging to the family Sittidae. Characterised by large heads, short tails, and powerful bills and feet, nuthatches advertise their territory using loud, simple songs. Mo ...
:
nuthatch The nuthatches () constitute a genus, ''Sitta'', of small passerine birds belonging to the family Sittidae. Characterised by large heads, short tails, and powerful bills and feet, nuthatches advertise their territory using loud, simple songs. M ...
es **
Tichodromidae The wallcreeper (''Tichodroma muraria'') is a small passerine bird found throughout the high mountains of the Palearctic from southern Europe to central China. It is the only extant member of both the genus ''Tichodroma'' and the family Tichodr ...
:
wallcreeper The wallcreeper (''Tichodroma muraria'') is a small passerine bird found throughout the high mountains of the Palearctic from southern Europe to central China. It is the only extant member of both the genus ''Tichodroma'' and the family Tichodr ...
**
Certhiidae The treecreepers are a family, Certhiidae, of small passerine birds, widespread in wooded regions of the Northern Hemisphere and sub-Saharan Africa. The family contains eleven species in two genera, ''Certhia'' and '' Salpornis''. Their plumage i ...
:
treecreeper The treecreepers are a family, Certhiidae, of small passerine birds, widespread in wooded regions of the Northern Hemisphere and sub-Saharan Africa. The family contains eleven species in two genera, '' Certhia'' and '' Salpornis''. Their plumage ...
s ** Mimidae:
mockingbird Mockingbirds are a group of New World passerine birds from the family Mimidae. They are best known for the habit of some species mimicking the songs of other birds and the sounds of insects and amphibians, often loudly and in rapid succession. ...
s and
thrasher Thrashers are a New World group of perching bird, passerine birds related to mockingbirds and New World catbirds. Like these, they are in the family (biology), family Mimidae. There are 15 species in one large and 4 monotypic genus, genera. T ...
s ** Sturnidae:
starling Starlings are small to medium-sized passerine birds in the family Sturnidae. The Sturnidae are named for the genus '' Sturnus'', which in turn comes from the Latin word for starling, ''sturnus''. Many Asian species, particularly the larger ones, ...
s ** Buphagidae:
oxpecker The oxpeckers are two species of bird which make up the genus ''Buphagus'', and family Buphagidae. The oxpeckers were formerly usually treated as a subfamily, Buphaginae, within the starling family, Sturnidae, but molecular phylogenetic studi ...
s **
Turdidae The thrushes are a passerine bird family, Turdidae, with a worldwide distribution. The family was once much larger before biologists reclassified the former subfamily Saxicolinae, which includes the chats and European robins, as Old World fl ...
: thrushes and allies **
Muscicapidae The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, Bluethroat (''Luscinia svecica)'' and Northe ...
:
Old World flycatcher The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, Bluethroat (''Luscinia svecica)'' and Northe ...
s and chats ** Cinclidae:
dipper Dippers are members of the genus ''Cinclus'' in the bird family Cinclidae, so-called because of their bobbing or dipping movements. They are unique among passerines for their ability to dive and swim underwater. Taxonomy The genus ''Cinclus'' ...
s ** Chloropseidae: leafbirds **
Dicaeidae The flowerpeckers are a family, Dicaeidae, of passerine birds. The family comprises two genera, '' Prionochilus'' and '' Dicaeum'', with 50 species in total. The family has sometimes been included in an enlarged sunbird family Nectariniidae. The ...
:
flowerpecker The flowerpeckers are a family, Dicaeidae, of passerine birds. The family comprises two genera, '' Prionochilus'' and '' Dicaeum'', with 50 species in total. The family has sometimes been included in an enlarged sunbird family Nectariniidae. The ...
s **
Nectariniidae Sunbirds and spiderhunters make up the family Nectariniidae of passerine birds. They are small, slender passerines from the Old World, usually with downward-curved bills. Many are brightly coloured, often with iridescent feathers, particularly ...
:
sunbird Sunbirds and spiderhunters make up the family Nectariniidae of passerine birds. They are small, slender passerines from the Old World, usually with downward-curved bills. Many are brightly coloured, often with iridescent feathers, particularly ...
s **
Passeridae Old World sparrows are a group of small passerine birds forming the family Passeridae. They are also known as true sparrows, a name also used for a particular genus of the family, '' Passer''. They are distinct from both the New World sparrows, ...
: true sparrows **
Ploceidae Ploceidae is a family of small passerine birds, many of which are called weavers, weaverbirds, weaver finches and bishops. These names come from the nests of intricately woven vegetation created by birds in this family. In most recent classifica ...
: weavers and widowbirds **
Estrildidae Estrildidae, or estrildid finches, is a family of small seed-eating passerine birds of the Old World tropics and Australasia. They comprise species commonly known as munias, mannikins, firefinches, parrotfinches and waxbills. Despite the word "fi ...
: estrildid finches ( waxbills, munias, etc.) **
Viduidae __NOTOC__ The indigobirds and whydahs, together with the cuckoo-finch, make up the family Viduidae; they are small passerine birds native to Africa. These are finch-like species which usually have black or indigo predominating in their plumage ...
: indigo birds and whydahs ** Peucedramidae:
olive warbler The olive warbler (''Peucedramus taeniatus'') is a small passerine bird. It is the only member of the genus ''Peucedramus'' and the family Peucedramidae. This species breeds from southern Arizona and New Mexico, USA, south through Mexico to Nica ...
**
Prunellidae The accentors are a genus of birds in the family Prunellidae, which is endemic to the Old World. This small group of closely related passerines are all in the genus ''Prunella''. All but the dunnock and the Japanese accentor are inhabitants of th ...
: accentor **
Motacillidae The wagtails, longclaws, and pipits are a family, Motacillidae, of small passerine birds with medium to long tails. Around 70 species occur in five genera. The longclaws are entirely restricted to the Afrotropics, and the wagtails are predomin ...
:
wagtail Wagtails are a group of passerine birds that form the genus ''Motacilla'' in the family Motacillidae. The forest wagtail belongs to the monotypic genus ''Dendronanthus'' which is closely related to ''Motacilla'' and sometimes included therein. ...
s and
pipit The pipits are a cosmopolitan genus, ''Anthus'', of small passerine birds with medium to long tails. Along with the wagtails and longclaws, the pipits make up the family Motacillidae. The genus is widespread, occurring across most of the world, ...
s ** Urocynchramidae: Przevalski's finch **
Fringillidae The true finches are small to medium-sized passerine birds in the family Fringillidae. Finches have stout conical bills adapted for eating seeds and nuts and often have colourful plumage. They occupy a great range of habitats where they are usua ...
: true
finch The true finches are small to medium-sized passerine birds in the family Fringillidae. Finches have stout conical bills adapted for eating seeds and nuts and often have colourful plumage. They occupy a great range of habitats where they are usua ...
es and
Hawaiian honeycreeper Hawaiian honeycreepers are a group of small, passerine birds endemic to Hawaii. They are closely related to the rosefinches in the genus ''Carpodacus'', but many species have evolved features unlike those present in any other finch. Their great ...
s (formerly Drepanididae) **
Parulidae The New World warblers or wood-warblers are a group of small, often colorful, passerine birds that make up the family Parulidae and are restricted to the New World. They are not closely related to Old World warblers or Australian warblers. Mos ...
: New World
warbler Various Passeriformes (perching birds) are commonly referred to as warblers. They are not necessarily closely related to one another, but share some characteristics, such as being fairly small, vocal, and insectivorous. Sylvioid warblers Th ...
s, for example the
black-throated blue warbler The black-throated blue warbler (''Setophaga caerulescens'') is a small passerine bird of the New World warbler family. Its breeding ranges are located in the interior of deciduous and mixed coniferous forests in eastern North America. Over the ...
s and allies **
Icteridae Icterids () or New World blackbirds make up a family, the Icteridae (), of small to medium-sized, often colorful, New World passerine birds. Most species have black as a predominant plumage color, often enlivened by yellow, orange, or red. Th ...
: American blackbirds,
New World oriole New World orioles are a group of birds in the genus ''Icterus'' of the blackbird family. Unrelated to Old World orioles of the family Oriolidae, they are strikingly similar in size, diet, behavior, and strongly contrasting plumage. As a resu ...
s,
grackle Grackles is the common name of any of 11 passerine birds (10 extant and one extinct) native to North and South America. They belong to various genera in the icterid family. In all the species with this name, adult males have black or mostly blac ...
s and
cowbird Cowbirds are birds belonging to the genus ''Molothrus'' in the family Icteridae. They are of New World origin, and are obligate brood parasites, laying their eggs in the nests of other species. The genus was introduced by English naturalist Will ...
s. ** Coerebidae: bananaquit **
Emberizidae The buntings are a group of Old World passerine birds forming the genus ''Emberiza'', the only genus in the family Emberizidae. The family contains 45 species. They are seed-eating birds with stubby, conical bills. Taxonomy The family Emberizid ...
: buntings **
Passerellidae New World sparrows are a group of mainly New World passerine birds, forming the family Passerellidae. They are seed-eating birds with conical bills, brown or gray in color, and many species have distinctive head patterns. Although they share t ...
:
New World sparrow New World sparrows are a group of mainly New World passerine birds, forming the family Passerellidae. They are seed-eating birds with conical bills, brown or gray in color, and many species have distinctive head patterns. Although they share t ...
s **
Thraupidae The tanagers (singular ) comprise the bird family Thraupidae, in the order Passeriformes. The family has a Neotropical distribution and is the second-largest family of birds. It represents about 4% of all avian species and 12% of the Neotropica ...
:
tanager The tanagers (singular ) comprise the bird family Thraupidae, in the order Passeriformes. The family has a Neotropical distribution and is the second-largest family of birds. It represents about 4% of all avian species and 12% of the Neotropica ...
s, true honeycreepers and allies **
Calcariidae Calcariidae is a small family of passerine birds. It includes longspurs and snow buntings. There are six species in three genera worldwide, found mainly in North America and Eurasia. They are migratory and can live in a variety of habitats includ ...
:
snow bunting The snow bunting (''Plectrophenax nivalis'') is a passerine bird in the family Calcariidae. It is an Arctic specialist, with a circumpolar Arctic breeding range throughout the northern hemisphere. There are small isolated populations on a few ...
s and
longspur The longspurs, genus ''Calcarius'', are a group of birds in the family Calcariidae. The name refers to the long claw on the hind toe of each foot. The genus formerly included the thick-billed longspur, ''Rhyncophanes mccownii'', which is now p ...
s **
Cardinalidae Cardinalidae (often referred to as the "cardinal-grosbeaks" or simply the "cardinals") is a family of New World-endemic passerine birds that consists of cardinals, grosbeaks, and buntings. It also includes several birds such as the tanager-lik ...
: cardinals and allies


See also

* Song system


References


Video links

* http://www.cbc.ca/natureofthings/blog/are-songbirds-disappearing
Mockingbird singing


External links


Oscines
Tree of Life web project article July 31, 2006. {{Authority control Extant Eocene first appearances