Licking
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Licking is the
action Action may refer to: * Action (philosophy), something which is done by a person * Action principles the heart of fundamental physics * Action (narrative), a literary mode * Action fiction, a type of genre fiction * Action game, a genre of video gam ...
of passing the
tongue The tongue is a Muscle, muscular organ (anatomy), organ in the mouth of a typical tetrapod. It manipulates food for chewing and swallowing as part of the digestive system, digestive process, and is the primary organ of taste. The tongue's upper s ...
over a surface, typically either to deposit
saliva Saliva (commonly referred as spit or drool) is an extracellular fluid produced and secreted by salivary glands in the mouth. In humans, saliva is around 99% water, plus electrolytes, mucus, white blood cells, epithelial cells (from which ...
onto the surface, or to collect liquid, food or minerals onto the tongue for
ingestion Ingestion is the consumption of a substance by an organism. In animals, it normally is accomplished by taking in a substance through the mouth into the gastrointestinal tract, such as through eating or drinking. In single-celled organisms, inge ...
, or to communicate with other animals. Many animals both groom themselves, eat or
drink A drink or beverage is a liquid intended for human consumption. In addition to their basic function of satisfying thirst, drinks play important roles in human culture. Common types of drinks include plain drinking water, milk, juice, smoothie ...
by licking.


In animals

Grooming: Animals commonly clean themselves through licking. In
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s, licking helps keep the
fur A fur is a soft, thick growth of hair that covers the skin of almost all mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an ...
clean and untangled. The tongues of many mammals have a rough upper surface that acts like a brush when the animal licks its fur. Certain reptiles, such as
gecko Geckos are small, mostly carnivorous lizards that have a wide distribution, found on every continent except Antarctica. Belonging to the infraorder Gekkota, geckos are found in warm climates. They range from . Geckos are unique among lizards ...
s, clean their
eye An eye is a sensory organ that allows an organism to perceive visual information. It detects light and converts it into electro-chemical impulses in neurons (neurones). It is part of an organism's visual system. In higher organisms, the ey ...
s by licking them, due to them not having eye lids. Mammals typically lick their offspring clean immediately after
birth Birth is the act or process of bearing or bringing forth offspring, also referred to in technical contexts as parturition. In mammals, the process is initiated by hormones which cause the muscular walls of the uterus to contract, expelling the f ...
; in many species this is necessary to free the newborn from the
amniotic sac The amniotic sac, also called the bag of waters or the membranes, is the sac in which the embryo and later fetus develops in amniotes. It is a thin but tough transparent pair of biological membrane, membranes that hold a developing embryo (and l ...
. The licking not only cleans and dries the offspring's fur, but also stimulates its
breath Breathing (spiration or ventilation) is the neuroscience of rhythm, rhythmical process of moving air into (inhalation) and out of (exhalation) the lungs to facilitate gas exchange with the Milieu intérieur, internal environment, mostly to flu ...
ing and digestive processes.
Canid Canidae (; from Latin, ''canis'', "dog") is a family (biology), biological family of caniform carnivorans, constituting a clade. A member of this family is also called a canid (). The family includes three subfamily, subfamilies: the Caninae, a ...
s also stimulate their pups to
urinate Urination is the release of urine from the bladder through the urethra in placental mammals, or through the cloaca in other vertebrates. It is the urinary system's form of excretion. It is also known medically as micturition, voiding, ure ...
by licking their preputial gland secretions. Food and water acquisition:
Hummingbird Hummingbirds are birds native to the Americas and comprise the Family (biology), biological family Trochilidae. With approximately 366 species and 113 genus, genera, they occur from Alaska to Tierra del Fuego, but most species are found in Cen ...
s are often said to "sip" nectar, but in fact they up nectar on their long tongues. Their tongues have fringed edges, which help both in nectar-eating and in catching tiny insects. Mother hummingbirds also lick their chicks after a rainstorm to dry them by licking water droplets from the coats of the chicks to avoid them chilling. Many animals drink by licking. While young mammals drink milk from their mothers' teats by
sucking Suction is the day-to-day term for the movement of gases or liquids along a pressure gradient with the implication that the movement occurs because the lower pressure pulls the gas or liquid. However, the forces acting in this case do not orig ...
, the typical method of drinking for adult mammals involves dipping the tongue repeatedly into water and using it to scoop water into the mouth. This method of drinking relies in part on the water adhering to the surface of the tongue and in part on muscular control of the tongue to form it into a
spoon A spoon (, ) is a utensil consisting of a shallow bowl (also known as a head), oval or round, at the end of a handle. A type of cutlery (sometimes called flatware in the United States), especially as part of a table setting, place setting, it ...
like shape. Cattle, horses and other animals lick rocks, salt licks or other objects to obtain mineral nutrients. Gustation: Animals also use their tongues to enhance their sense of smell. By licking a surface or extending the tongue beyond the mouth, molecules are transferred via the tongue to the olfactory receptors in the
nose A nose is a sensory organ and respiratory structure in vertebrates. It consists of a nasal cavity inside the head, and an external nose on the face. The external nose houses the nostrils, or nares, a pair of tubes providing airflow through the ...
and in some animals, to the
vomeronasal organ The vomeronasal organ (VNO), or Jacobson's organ, is the paired auxiliary olfactory (smell) sense organ located in the soft tissue of the nasal septum, in the nasal cavity just above the roof of the mouth (the hard palate) in various tetrapods ...
. In some mammals, the tongue is used to "lick" the air during the flehmen response to assist transfer of pheremones. Similarly, snakes use smell to track their prey. They smell by using their forked tongues to collect airborne particles, then passing them to the vomeronasal organ. They keep their tongues constantly in motion, sampling particles from the air, ground, and water, analyzing the chemicals found, and determining the presence of prey or predators in the local environment. Communication: Dogs and cats use licking both to clean and to show affection among themselves or to humans, typically licking their faces. Many animals use licking as a submissive or appeasement signal in dominance hierarchies. Thermoregulation: Some animals use licking to cool themselves. Cats do not
sweat Perspiration, also known as sweat, is the fluid secreted by sweat glands in the skin of mammals. Two types of sweat glands can be found in humans: eccrine glands and Apocrine sweat gland, apocrine glands. The eccrine sweat glands are distribu ...
the way humans do and the saliva deposited by licking provides a similar means of
evaporative cooling An evaporative cooler (also known as evaporative air conditioner, swamp cooler, swamp box, desert cooler and wet air cooler) is a device that cools air through the evaporation of water. Evaporative cooling differs from other air conditioning sy ...
. Some animals spread saliva over areas of the body with little or no fur to maximise heat loss. For example,
kangaroo Kangaroos are marsupials from the family Macropodidae (macropods, meaning "large foot"). In common use, the term is used to describe the largest species from this family, the red kangaroo, as well as the antilopine kangaroo, eastern gre ...
s lick their wrists and
rat Rats are various medium-sized, long-tailed rodents. Species of rats are found throughout the order Rodentia, but stereotypical rats are found in the genus ''Rattus''. Other rat genera include '' Neotoma'' (pack rats), '' Bandicota'' (bandicoo ...
s lick their testicles. Mating behavior: Male mammals often lick the genitals of females before copulation. Post-copulatory genital grooming often occurs in male rats and
prosimian Prosimians are a group of primates that includes all living and extinct Strepsirrhini, strepsirrhines (lemurs, Lorisoidea, lorisoids, and Adapiformes, adapiforms), as well as the Haplorhini, haplorhine tarsiers and their extinct relatives, the Om ...
primates. This behavior may prevent disease transmission.


In primates

Ring-tailed lemurs lick each other's babies as a means of collective grooming and of reinforcing social cohesion within the community. Macaques and other primates lick leaves for water in addition to dipping their arms into tree crevices and licking the water off. Chimpanzees use licking in a variety of ways: licking objects, such as dead trees, that others in their community have licked,楽天が運営するポータルサイト : 【インフォシーク】Infoseek
. Mahale.web.infoseek.co.jp (2000-01-01). Retrieved on 2013-03-30.
licking each other's body parts for grooming and sex and licking rocks for salt. Gorillas use licking in addition to other senses to determine the nature of an object.


In humans

Compared to most other mammals, licking has a minor role for
human Humans (''Homo sapiens'') or modern humans are the most common and widespread species of primate, and the last surviving species of the genus ''Homo''. They are Hominidae, great apes characterized by their Prehistory of nakedness and clothing ...
s. The human tongue is relatively short and inflexible, and is not well adapted for either grooming or drinking. Instead, humans prefer to wash themselves using their hands and drink by
sucking Suction is the day-to-day term for the movement of gases or liquids along a pressure gradient with the implication that the movement occurs because the lower pressure pulls the gas or liquid. However, the forces acting in this case do not orig ...
or pouring fluid into their mouth. Humans have much less hair over their skin than most other mammals, and much of that hair is in places which they cannot reach with their own mouth. The presence of
sweat glands Sweat glands, also known as sudoriferous or sudoriparous glands, , are small tubular structures of the skin that produce sweat. Sweat glands are a type of exocrine gland, which are glands that produce and secrete substances onto an epithelial sur ...
all over the human body makes licking as a cooling method unnecessary. Nonetheless, licking does play a role for humans. Even though humans cannot effectively drink water by licking, the human tongue is quite sufficient for licking more
viscous Viscosity is a measure of a fluid's rate-dependent resistance to a change in shape or to movement of its neighboring portions relative to one another. For liquids, it corresponds to the informal concept of ''thickness''; for example, syrup h ...
fluids. Some foods are sold in a form intended to be consumed mainly by licking, e.g.
ice cream Ice cream is a frozen dessert typically made from milk or cream that has been flavoured with a sweetener, either sugar or an alternative, and a spice, such as Chocolate, cocoa or vanilla, or with fruit, such as strawberries or peaches. Food ...
cones and
lollipop A lollipop is a type of sugar candy usually consisting of hard candy mounted on a stick and intended for sucking or licking. Different informal terms are used in different places, including lolly, sucker and sticky-pop. Lollipops are avail ...
s. Though useful, in some cultures it is considered improper table manners to clean one's fingers by licking during a meal. Some people in the Afar tribe of Ethiopia have been reported to have used their tongues to lick other humans, as a way of cleaning them from the dust that accumulates on them in a very water-scarce region. Humans use licking for a number of other purposes. For example, licking can moisten the
adhesive Adhesive, also known as glue, cement, mucilage, or paste, is any non-metallic substance applied to one or both surfaces of two separate items that binds them together and resists their separation. The use of adhesives offers certain advantage ...
surfaces of stamps or
envelope An envelope is a common packaging item, usually made of thin, flat material. It is designed to contain a flat object, such as a letter (message), letter or Greeting card, card. Traditional envelopes are made from sheets of paper cut to one o ...
s. Many people lick a fingertip (usually that of the
index finger The index finger (also referred to as forefinger, first finger, second finger, pointer finger, trigger finger, digitus secundus, digitus II, and many other terms) is the second digit of a human hand. It is located between the thumb and the m ...
) for some extra grip when turning a page, taking a sheet of paper from the top of a pile or opening a plastic bag. In
sewing Sewing is the craft of fastening pieces of textiles together using a sewing needle and thread. Sewing is one of the oldest of the textile arts, arising in the Paleolithic era. Before the invention of spinning yarn or weaving fabric, archaeo ...
, thread ends are commonly wet by licking to make the
fibre Fiber (spelled fibre in British English; from ) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often incorp ...
s stick together and thus make threading them through the eye of a needle easier. Another practice considered uncivilized is licking one's hand and using it to groom one's hair. Humans also use their tongues for sexual purposes, such as during
cunnilingus Cunnilingus is an oral sex act consisting of the stimulation of a vulva by using the tongue and lips. The clitoris is the most sexually sensitive part of the vulva, and its stimulation may result in a woman becoming sexually aroused or achievi ...
,
anilingus Anilingus (also spelled analingus) is an oral sex, oral and anal sex act (anal–oral contact or anal–oral sex) in which one person stimulates the human anus, anus of another by using their tongue or lips. The anus has a relatively high conc ...
,
fellatio Fellatio (also known as fellation, and in slang as blowjob, BJ, giving head, or sucking off) is an oral sex act consisting of the stimulation of a human penis, penis by using the mouth. Oral stimulation of the scrotum may also be termed ''fellat ...
, breast licking, foot licking, and whilst
French kiss A French kiss, also known as cataglottism or a tongue kiss, is an amorous kiss in which the participants' tongues extend to touch each other's lips or tongue. A kiss with the tongue stimulates the partner's lips, tongue and mouth, which are sens ...
ing, where two people lick each other's tongues.


In dogs


Abnormal licking

Self-licking can sometimes become abnormally frequent occasionally resulting in a
lick granuloma A lick granuloma, also known as acral lick dermatitis, is a skin disorder found most commonly in dogs, but also in cats. In dogs, it results typically from the dog's urge to lick the lower portion of one of their legs. The lesion can initially be ...
. The most common cause of lick granuloma appears to be psychological, related to stress, anxiety,
separation anxiety Separation anxiety disorder (SAD) is an anxiety disorder in which an individual experiences excessive anxiety regarding separation from home and/or from people to whom the individual has a strong emotional attachment (e.g., a parent, caregive ...
, boredom, or compulsiveness. Lick granulomae are especially seen in active dogs left alone for long periods of time. One theory is that excessive licking causes
endorphin Endorphins (contracted from endogenous morphine) are peptides produced in the brain that block the perception of pain and increase feelings of wellbeing. They are produced and stored in the pituitary gland of the brain. Endorphins are endogeno ...
release, which reduces pain and makes the dog feel temporarily
euphoric Euphoria ( ) is the experience (or affect) of pleasure or excitement and intense feelings of well-being and happiness. Certain natural rewards and social activities, such as aerobic exercise, laughter, listening to or making music and danci ...
. This provides the animal with positive feedback from the licking, and subsequent addiction to the behaviour. Animals in captivity sometimes develop a licking
stereotypy A stereotypy (, ) is a repetitive or ritualistic movement, posture, or utterance. Stereotypies may be simple movements such as body rocking, or complex, such as self-caressing, crossing and uncrossing of legs, and marching in place. They are foun ...
during which surfaces (walls, bars, gates, etc.) are repeatedly licked for no apparent reason. This has been observed in captive giraffes and camels.Video of a giraffe licking wall


References

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External links


Grooming in cats

Hummingbird Licking Behavior

Mice Licking Responses to Sweeteners



Video of a giraffe licking a wall
Eating behaviors Ethology Saliva Tongue Hygiene Animal communication Mating