''Hsanotherium'' is an extinct genus of early ungulate from the middle

Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...

, described in 2000 in the Pondaung Formation,

Myanmar Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and has ...

. It was originally assigned to

Anthracobunidae Anthracobunidae is an extinct family of stem perissodactyls that lived in the early to middle Eocene period. They were originally considered to be a paraphyletic family of primitive proboscideans possibly ancestral to the Moeritheriidae and th ...

(formerly considered

proboscidea Proboscidea (; , ) is a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Three l ...

ns, now

perissodactyl Perissodactyla (, ), or odd-toed ungulates, is an order of Ungulate, ungulates. The order includes about 17 living species divided into three Family (biology), families: Equidae (wild horse, horses, Asinus, asses, and zebras), Rhinocerotidae ( ...

s) making it the first of that family to be discovered outside India and Pakistan and also the smallest. However, the authors of a 2014 cladistic study regard it as more similar to medium-sized bunodont

artiodactyl Artiodactyls are placental mammals belonging to the order Artiodactyla ( , ). Typically, they are ungulates which bear weight equally on two (an even number) of their five toes (the third and fourth, often in the form of a hoof). The other t ...

s, such as '' Haqueina''.

References

* DUCROCQ S.; AUNG NAING SOE; BO B.; BENAMMI M.; CHAIMANEE Y.; TUN T.; THEINE T.; JAEGER J.-J.; First record of an Anthracobunidae (Mammalia, ?Tethytheria) from the Eocene of the Pondaung Formation, Myanmar, Comptes rendus de l'Académie des sciences. Série 2. Sciences de la terre et des planètes (C. r. Acad. sci., Sér. 2, Sci. terre planet.), 2000, vol. 330, no10, pp. 725–730 Fossil taxa described in 2000 Eocene mammals of Asia Prehistoric Perissodactyla {{paleo-oddtoedungulate-stub