The Satyrini is one of the

tribes The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...

of the

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

Satyrinae The Satyrinae, the satyrines or satyrids, commonly known as the browns, are a subfamily of the Nymphalidae (brush-footed butterflies). They were formerly considered a distinct family, Satyridae. This group contains nearly half of the known divers ...

. It includes about 2200

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

and is therefore the largest tribe in the subfamily which comprises 2500 species.

Distribution

Satyrini butterflies have a worldwide distribution, but the distribution pattern differs between

subtribe Subtribe is a taxonomic category ranking which is below the rank of tribe and above genus. The standard suffix for a subtribe is -ina (in animals) or -inae (in plants). The first use of this word dates back to the late 19th century. An example of ...

s. Some subtribes are almost restricted to a single

biogeographic region An ecoregion (ecological region) is an ecological and geographic area that exists on multiple different levels, defined by type, quality, and quantity of environmental resources. Ecoregions cover relatively large areas of land or water, and con ...

, such as the

Pronophilina Pronophilina is a Neotropical subtribe of butterflies of the subfamily Satyrinae. They are a species-rich group with highest diversity in the tropical and subtropical mountains, especially the Andes. Before 1970, they were poorly studied, but rec ...

, which is found only in

Andean The Andes ( ), Andes Mountains or Andean Mountain Range (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long and wide (widest between 18°S ...

cloud forests A cloud forest, also called a water forest, primas forest, or tropical montane cloud forest, is a generally tropical or subtropical, evergreen, montane, moist forest characterized by a persistent, frequent or seasonal low-level cloud cover, ...

from

Venezuela Venezuela, officially the Bolivarian Republic of Venezuela, is a country on the northern coast of South America, consisting of a continental landmass and many Federal Dependencies of Venezuela, islands and islets in the Caribbean Sea. It com ...

Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...

Biology

The

larval food plants A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...

of many species in this tribe are

grasses Poaceae ( ), also called Gramineae ( ), is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos, the grasses of natural grassland and species cultivated in ...

, i.e. Poaceae. It is considered that the Satyrini diversified at about the same time as the grasses did, and that the

radiation In physics, radiation is the emission or transmission of energy in the form of waves or particles through space or a material medium. This includes: * ''electromagnetic radiation'' consisting of photons, such as radio waves, microwaves, infr ...

of the tribe is therefore closely related to the evolution of the grasses. In contrast, the tribe has a few genera which show uncommon feeding preferences. Three genera, ''

Euptychia ''Euptychia'' is a genus of satyrid butterflies found in the Neotropical realm. The genus was erected by Jacob Hübner Jacob Hübner (20 June 1761 – 13 September 1826, in Augsburg) was a German entomologist. He was the author of ''Sammlung E ...

'', ''

Ragadia ''Ragadia'' is a genus of brush-footed butterflies (family Nymphalidae The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to the superfamily Papilion ...

'' and ''

Acrophtalmia Acrophtalmia is a genus of butterflies in the family Nymphalidae The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to the superfamily Papilionoidea, ...

'', feed on

Lycopsida Lycopodiopsida is a class of vascular plants also known as lycopsids, lycopods, or lycophytes. Members of the class are also called clubmosses, firmosses, spikemosses and quillworts. They have dichotomously branching stems bearing simple leaves ...

, and moreover, some species of ''Euptychia'' have been reported to feed on

mosses Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta ('' sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and ho ...

Neckeraceae Neckeraceae is a moss family in the order Hypnales. There are about 200 species native to temperate and tropical regions. Most grow on rocks, or other plants. Description Members of the family are usually large and glossy plants with creeping st ...

. This is interesting cases as these genera are not considered to be very closely related to each other.

Taxonomy

The taxonomy of the subfamily Satyrinae has recently undergone a significant revision. showed that most of the traditionally well known group are

paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...

polyphyletic A polyphyletic group is an assemblage that includes organisms with mixed evolutionary origin but does not include their most recent common ancestor. The term is often applied to groups that share similar features known as Homoplasy, homoplasies ...

and developed a new taxonomy, based on

molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

. The classification scheme transferred many genera traditionally placed in the

Elymniini The Elymniini is one of the tribe (biology), tribes of the subfamily Satyrinae. If the subfamily Satyrinae elevate to family (biology), family status, this tribe shall be treated as subfamily Elymniinae. Elymniini was formerly a large group, but ...

to the Satyrini, and later studies have largely followed this classification. Thus, the Satyrini is now considered to include the genera previously placed in the Elymniini, such as ''

Lethe In Greek mythology, Lethe (; Ancient Greek: ''Lḗthē''; , ) was one of the rivers of the underworld of Hades. In Classical Greek, the word '' lethe'' ( λήθη) literally means "forgetting", "forgetfulness". The river is also known as Amel ...

'', '' Pararge'' and ''

Mycalesis ''Mycalesis'', the bushbrowns, are a genus of Nymphalidae, brush-footed butterfly, butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymo ...

''. However, the phylogenetic statuses of some subtribes within the tribe are still unclear and the revision of the classification is likely to continue. ---- According to , the Satyrini includes the following

s and

genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...

: ; Parargina Moore, 1893 :* '' Chonala'' Moore, 1893 :* '' Kirinia'' Westwood, 1841 :* ''

Lasiommata ''Lasiommata'' is a genus of butterflies of the subfamily Satyrinae in the family Nymphalidae. Species * ''Lasiommata megera'' (Linnaeus, 1767)Lopinga ''Lopinga'' is a genus of butterflies of the family Nymphalidae. ''Lopinga achine,'' a threatened species classified as vulnerable in Swedish Red data book (Ehnström et al., 1993) This butterfly frequents glades in partly open oak woodland wher ...

'' Moore, 1893 :* '' Orinoma'' Gray, 1846 :* '' Pararge'' Hübner, 1819 :* '' Rhaphicera'' Butler, 1867 :* '' Tatinga'' Moore, 1893 :* '' Nosea'' Koiwaya 1993 ;

Lethina The Satyrini is one of the tribe (biology), tribes of the subfamily Satyrinae. It includes about 2200 species and is therefore the largest tribe in the subfamily which comprises 2500 species. Distribution Satyrini butterflies have a Cosmopolit ...

Reuter, 1896 :* ''

Aphysoneura ''Aphysoneura'' is a genus of butterflies from the subfamily Satyrinae in the family Nymphalidae The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to ...

'' Karsch, 1894 :* ''

'' Hübner, 1819 :* '' Neope'' Moore, 1866 :* ''

Ninguta Ning'an () is a city located approximately southwest of Mudanjiang, in the southeast of Heilongjiang province, China, bordering Jilin province to the south. It is located on the Mudanjiang River (formerly known as Hurka River), which flows north, ...

'' Moore, 1892 :* '' Ptychandra'' Felder & Felder, 1861 :* '' Mandarinia'' Leech, 1892 ;

Mycalesina The Satyrini is one of the tribes of the subfamily Satyrinae. It includes about 2200 species and is therefore the largest tribe in the subfamily which comprises 2500 species. Distribution Satyrini butterflies have a worldwide distribution, bu ...

Reuter, 1896 :* '' Lohora'' Moore, 1880 :* '' Brakefieldia'' Aduse-Poku, Lees & Wahlberg, 2016 :* '' Telinga'' Moore, 1880 :* '' Heteropsis'' Westwood, 1850 :* '' Culapa'' Moore, 1879 :* '' Mydosama'' Moore, 1880 :* ''

'' Hübner, 1818 :* '' Devyatkinia'' Monastyrskii & Uémura, 2016 :* '' Hallelesis'' Condamin, 1961 :* ''

Bicyclus ''Bicyclus'' is a butterfly genus from the subfamily Satyrinae in the family Nymphalidae. The species are found in the Afrotropical realm. Species *''Bicyclus abnormis'' (Dudgeon, 1909) *''Bicyclus albocincta'' (Rebel, 1914) *''Bicyclus alboplag ...

'' Kirby, 1871 ; Eritina Miller, 1968 :* '' Coelites'' Westwood, 1850 :* '' Erites'' Westwood, 1851 :* '' Orsotriaena'' Wallengren, 1858 :* '' Zipaetis'' Hewitson, 1863 ; Ragadiina Herrich-Schäffer, 1864 :* ''

'' Felder & Felder, 1861 :* ''

Acropolis An acropolis was the settlement of an upper part of an ancient Greek city, especially a citadel, and frequently a hill with precipitous sides, mainly chosen for purposes of defense. The term is typically used to refer to the Acropolis of Athens ...

'' Hemming, 1934 :* ''

'' Westwood, 1851 ; Coenonymphina Tutt, 1896 :* ''

Coenonympha ''Coenonympha'' is a butterfly genus belonging to the Coenonymphina, a subtribe of the browns (Satyrinae). The latter are a subfamily of the brush-footed butterflies (Nymphalidae). As a rule, Palearctic species are colloquially called heaths, whi ...

'' Hübner, 1819 :* '' Sinonympha'' Lee, 1994 :* '' Oressinoma'' Doubleday, 1849 :* '' Altiapa'' Parsons, 1986 :* ''

Argynnina Argynnini is a tribe of butterflies in the subfamily Heliconiinae, containing some of the fritillaries. This group has roughly 100 species worldwide and roughly 30 in North America. Systematics This group has also been classified as subtribe Ar ...

'' Butler, 1867 :* '' Argyronympha'' Mathew, 1886 :* '' Argyrophenga'' Doubleday, 1845 :* ''

Dodonidia The forest ringlet (''Dodonidia helmsii''), also known as Helms' butterfly, or te pēpepe pōuri or pepe pouri in the Māori language, is a rare butterfly of the family Nymphalidae endemic to New Zealand. It is the only species in the genus ''Do ...

'' Butler, 1884 :* '' Erebiola'' Fereday, 1879 :* '' Erycinidia'' Rothschild & Jordan, 1905 :* '' Geitoneura'' Butler, 1867 :* '' Harsiesis'' Fruhstorfer, 1911 :* '' Heteronympha'' Wallengren, 1858 :* '' Hyalodia'' Jordan, 1924 :* '' Hypocysta'' Westwood, 1851 :* '' Lamprolenis'' Godman & Salvin, 1881 :* '' Nesoxenica'' Waterhouse & Lyell, 1914 :* '' Oreixenica'' Waterhouse & Lyell, 1914 :* '' Paratisiphone'' Watkins, 1928 :* '' Percnodaimon'' Butler, 1876 :* '' Platypthima'' Rothschild & Jordan, 1905 :* ''

Tisiphone Tisiphone (, "Avenger of murder"),Tisiphone
Encyclopedia Britannica, Retrieved 4 February 2025; from '' ...

'' Hübner, 1819 ;

Euptychiina The butterfly subtribe Euptychiina (Lepidoptera: Nymphalidae) is a diverse group within the Tribe (biology), tribe Satyrini, occurring throughout Central America, Central and South America, in addition to a few species known from North America. E ...

Reuter, 1896 :* ''

'' Hübner, 1818 :* '' Palaeonympha'' Butler, 1871 :* ''

Amphidecta ''Amphidecta'' is a genus of satyrid butterfly found in the Neotropical realm The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropics, tropical Ecoregion#Terrestr ...

'' Butler, 1867 :* '' Archeuptychia'' Forster, 1964 :* ''

Amiga Amiga is a family of personal computers produced by Commodore International, Commodore from 1985 until the company's bankruptcy in 1994, with production by others afterward. The original model is one of a number of mid-1980s computers with 16-b ...

'' Nakahara, Willmott & Espeland, 2019 :* '' Caeruleuptychia'' Forster, 1964 :* '' Capronnieria'' Forster, 1964 :* '' Cepheuptychia'' Forster, 1964 :* '' Chloreuptychia'' Forster, 1964 :* '' Cissia'' Doubleday, 1848 :* '' Coeruleotaygetis'' Forster, 1964 :* ''

Cyllopsis ''Cyllopsis'' is a genus of satyrid butterflies found in the Neotropical realm. The genus was erected by Rudolf Felder in 1869. Species Listed alphabetically: *''Cyllopsis argentella'' (Butler & H. Druce, 1872) *''Cyllopsis caballeroi'' Beutels ...

'' Felder, 1869 :* '' Erichthodes'' Forster, 1964 :* '' Euptychoides'' Forster, 1964 :* ''

Forsterinaria ''Forsterinaria'' is a genus of satyrid butterflies found in the Neotropical realm. Species Listed alphabetically:Graphita'' Nakahara, Marín & Barbosa 2016 :* '' Godartiana'' Forster, 1964 :* '' Harjesia'' Forster, 1964 :* ''

Hermeuptychia ''Hermeuptychia'' is a genus of Satyrinae, satyrid butterflies found in the Neotropical realm. They are a widespread, cryptic genus, with Cytochrome c oxidase subunit I species delineation methods implying much greater species diversity than curr ...

'' Forster, 1964 :* '' Huberonympha'' Viloria & Costa, 2016 :* '' Inbio'' Nakahara & Espeland, 2015 :* '' Magneuptychia'' Forster, 1964 :* '' Megeuptychia'' Forster, 1964 :* '' Megisto'' Hübner, 1819 :* '' Moneuptychia'' Forsterm, 1964 :* ''

Neonympha ''Neonympha'' is a Nearctic and Neotropical genus of satyrid butterflies. Species Listed alphabetically: *'' Neonympha areolatus'' (Smith, 1797) *'' Neonympha helicta'' (Hübner, 808 808 may refer to: Music * Roland-808, Roland TR-808, a drum ...

'' Hübner 1818 :* '' Nhambikuara'' Freitas, Barbosa & Zacca, 2018 :* '' Optimandes'' Marín, Nakahara & Willmott, 2019 :* '' Orotaygetis'' Nakahara & Zacca, 2018 :* '' Paramacera'' Butler, 1868 :* '' Parataygetis'' Forster, 1964 :* '' Pareuptychia'' Forster, 1964 :* '' Paryphthimoides'' Forster, 1964 :* '' Pharneuptychia'' Forster, 1964 :* ''

Pindis ''Pindis'' is a genus of satyrid butterflies found in the Neotropical realm The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropics, tropical Ecoregion#Terrestria ...

'' Felder, 1869 :* '' Posttaygetis'' Forster, 1964 :* '' Prenda'' Freitas & Mielke, 2011 :* '' Pseudeuptychia'' Forster, 1964 :* '' Pseudodebis'' Forster, 1964 :* '' Rareuptychia'' Forster, 1964 :* '' Satyrotaygetis'' Forster, 1964 :* '' Sepona'' Freitas & Barbosa, 2016 :* '' Splendeuptychia'' Forster, 1964 :* '' Stegosatyrus'' Zacca, Mielke & Pyrcz, 2013 :* '' Stevenaria'' Viloria, Costa, Neild & Nakahara, 2016 :* '' Taydebis'' Freitas, 2003 :* '' Taygetina'' Forster, 1964 :* ''

Taygetis ''Taygetis'' is a genus of Satyrinae, satyrid butterflies found in the Neotropical realm. Taxonomy and evolution The genus appeared as polyphyletic,, 2013: Systematics and evolutionary history of butterflies in the “''Taygetis'' clade” (Nymp ...

'' Hübner, 1819 :* '' Taygetomorpha'' Miller, 2004 :* '' Yphthimoides'' Forster, 1964 :* '' Zischkaia'' Forster, 1964 ; Erebiina Tutt, 1896 :* ''

Erebia ''Erebia'' is a Holarctic genus of brush-footed butterflies, family (biology), family Nymphalidae. Most of the about 90–100 species (see also #Taxonomy and systematics, below) are dark brown or black in color, with reddish-brown to orang ...

'' Elwes, 1899 ; Maniolina Grote, 1897 :* ''

Aphantopus ''Aphantopus'' is a butterfly genus of the Satyrinae. The genus is confined to the Palearctic. Species *''Aphantopus arvensis'' (Oberthür, 1876) (western China) **''Aphantopus arvensis arvensis'' **''Aphantopus arvensis campana'' Leech, 1892 (c ...

'' Wallengren, 1853 :* '' Cercyonis'' Scudder, 1875 :* ''

Hyponephele ''Hyponephele'' is a genus of butterflies of the subfamily Satyrinae in the family Nymphalidae. Most range from Europe into central Asia with a few in northern Africa. They are commonly called meadowbrowns. Species Listed alphabetically:< ...

'' Muschamp, 1915 :* ''

Maniola ''Maniola'' is a genus of butterflies within the family Nymphalidae The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to the superfamily Papilionoid ...

'' Schrank, 1801 :* ''

Pyronia ''Pyronia'' is a genus of butterflies from the subfamily Satyrinae in the family Nymphalidae. Description These butterflies have vestigial Vestigiality is the retention, during the process of evolution, of genetically determined structure ...

'' Hübner, 1819 ; Melanargiina Wheeler, 1903 :* '' Melanargia'' Meigen, 1828 ;

Reuter, 1896 :* '' Altopedaliodes'' Forster, 1964 :* '' Antopedaliodes'' Forster, 1964 :* '' Apexacuta'' Pyrcz, 2004 :* ''

Arhuaco The Arhuaco are an Indigenous people of Colombia. They are Chibchan-speaking people and descendants of the Tairona culture, concentrated in northern Colombia in the Sierra Nevada de Santa Marta. Name The Arhuaco are also known as the Aruaco, ...

'' Adams & Bernard, 1977 :* '' Cheimas'' Thieme, 1907 :* '' Corades'' Doubleday, 1849 :* '' Corderopedaliodes'' Forster, 1964 :* '' Daedalma'' Hewitson, 1858 :* '' Dangond'' Adams & Bernard, 1979 :* '' Drucina'' Butler, 1872 :* '' Druphila'' Pyrcz, 2004 :* ''

Eretris ''Eretris'' is a Neotropical butterfly genus in the family Nymphalidae. The genus was erected by Theodor Otto Thieme in 1905. Species *''Eretris apuleina'' Pyrcz, 2004 *''Eretris apuleja'' (C. Felder & R. Felder, 1867) *''Eretris calisto'' (C. ...

'' Thieme, 1905 :* '' Eteona'' Doubleday, 1848 :* '' Foetterleia'' Viloria, 2004 :* '' Junea'' Hemming, 1964 :* '' Lasiophila'' Felder & Felder, 1859 :* '' Lymanopoda'' Westwood, 1851 :* '' Mygona'' Thieme, 1907 :* '' Neopedaliodes'' Viloria, Miller & Miller 2004 :* '' Oxeoschistus'' Butler, 1867 :* '' Panyapedaliodes'' Forster, 1964 :* '' Paramo'' Adams & Bernard, 1977 :* '' Parapedaliodes'' Forster, 1964 :* '' Pedaliodes'' Butler, 1867 :* '' Pherepedaliodes'' Forster, 1964 :* '' Physcopedaliodes'' Forster, 1964 :* '' Praepedaliodes'' Forster, 1964 :* '' Praepronophila'' Forster, 1964 :* ''

Proboscis A proboscis () is an elongated appendage from the head of an animal, either a vertebrate or an invertebrate. In invertebrates, the term usually refers to tubular arthropod mouthparts, mouthparts used for feeding and sucking. In vertebrates, a pr ...

'' Thieme, 1907 :* '' Pronophila'' Doubleday, 1849 :* '' Protopedaliodes'' Viloria & Pyrcz, 1994 :* '' Pseudomaniola'' Röber, 1889 :* '' Punapedaliodes'' Forster, 1964 :* '' Steremnia'' Thieme, 1905 :* '' Steroma'' Westwood, 1850 :* '' Steromapedaliodes'' Forster, 1964 :* '' Thiemeia'' Weymer, 1912 :* '' Diaphanos'' Adams & Bernard, 1981 :* '' Ianussiusa'' Pyrcz & Viloria, 2004 :* '' Idioneurula'' Strand, 1932 :* '' Manerebia'' Staudinger, 1897 :* '' Neomaniola'' Hayward, 1949 :* '' Argyrophorus'' Blanchard, 1852 :* '' Auca'' Hayward, 1953 :* '' Chillanella'' Herrera, 1966 :* '' Cosmosatyrus'' Felder & Felder, 1867 :* '' Elina'' Blanchard, 1852 :* ''

Faunula ''Faunula'' is a Neotropical genus of butterflies in the nymphalid subfamily Satyrinae The Satyrinae, the satyrines or satyrids, commonly known as the browns, are a subfamily of the Nymphalidae (brush-footed butterflies). They were formerly co ...

'' Felder & Felder, 1867 :* '' Haywardella'' Herrera, 1966 :* '' Homoeonympha'' Felder & Felder, 1867 :* ''

Nelia Nelia or Neleia ( or Νήλεια) was a town of Magnesia in ancient Thessaly; Demetrias was situated between it and Iolcus. Strabo Strabo''Strabo'' (meaning "squinty", as in strabismus) was a term employed by the Romans for anyone whose eye ...

'' Hayward, 1953 :* '' Neomaenas'' Wallengren, 1858 :* '' Neosatyrus'' Wallengren, 1858 :* '' Pampasatyrus'' Hayward, 1953 :* '' Quilaphoestosus'' Herrera, 1966 :* '' Spinantenna'' Hayward, 1953 :* '' Tetraphlebia'' Felder & Felder, 1867 :* '' Gyrocheilus'' Butler, 1867 ; Satyrina Boisduval, 1833 :* '' Argestina'' Riley, 1923 :* ''

Davidina ''Davidina '' is a genus of butterflies in the family Nymphalidae (subfamily Satyrinae). (central China: Shaanxi) References External linksImages representing ''Davidina''at Consortium for the Barcode of Life The Consortium for the Barcode o ...

'' Oberthür, 1879 :* '' Berberia'' de Lesse, 1951 :* '' Hipparchia'' Fabricius, 1807 :* '' Kanetisa'' Moore, 1893 :* '' Aulocera'' Butler, 1867 :* '' Karanasa'' Moore, 1893 :* ''

Oeneis ''Oeneis'' (the Arctics or graylings) is a butterfly genus of the Satyrinae. All but one of its members are Arctic, sub-Arctic or high-altitude alpine in distribution. Some of the members of the genus are among the butterflies that can get along ...

'' Hübner, 1819 :* '' Paroeneis'' Moore, 1893 :* '' Arethusana'' de Lesse, 1951 :* '' Brintesia'' Fruhstorfer, 1911 :* '' Minois'' Hübner, 1819 :* '' Chazara'' Moore, 1893 :* ''

Pseudochazara ''Pseudochazara'' is a genus of butterfly, butterflies within the family Nymphalidae. Species * ''Pseudochazara alpina'' (Staudinger, 1878) * ''Pseudochazara amalthea'' (Frivaldsky, 1845) * ''Pseudochazara anthelea'' (Hübner, 823-1824 * '' ...

'' de Lesse, 1951 :* '' Satyrus'' Latreille, 1810 ; Ypthimina Reuter, 1896 :* '' Cassionympha'' van Son, 1955 :* '' Coenyra'' Hewitson, 1865 :* '' Coenyropsis'' van Son, 1958 :* '' Mashuna'' van Son, 1955 :* '' Mashunoides'' Mendes & Bivar de Sousa, 2009 :* '' Melampias'' Hübner, 1819 :* '' Neita'' van Son, 1955 :* ''

Neocoenyra ''Neocoenyra'' is a genus of Afrotropical realm, Afrotropical butterflies from the subfamily Satyrinae in the family Nymphalidae. Species *''Neocoenyra bioculata'' Carcasson, 1964 *''Neocoenyra cooksoni'' Druce, 1907 *''Neocoenyra duplex'' Bu ...

'' Butler, 1886 :* ''

Pseudonympha ''Pseudonympha'' is a genus of butterflies from the subfamily Satyrinae in the family Nymphalidae.

'' Wallengren, 1857 :* '' Paternympha'' Henning & Henning, 1997 :* '' Physcaeneura'' Wallengren, 1857 :* ''

Strabena ''Strabena'' is an Afrotropical realm, Afrotropical butterfly genus Endemic species, endemic to Geography of Madagascar, Madagascar from the subfamily Satyrinae in the family Nymphalidae.Stygionympha'' van Son, 1955 :* ''

Ypthima ''Ypthima'' is a species-rich genus of Old World butterfly, butterflies in the family Nymphalidae.

'' Hübner, 1818 :* '' Austroypthima'' Holloway, 1974 :* '' Ypthimomorpha'' van Son, 1955 ; subtribe uncertain :* '' Calisto'' Hübner, 1823 :* '' Paralasa'' Moore, 1893 :* '' Callerebia'' Butler, 1867 :* '' Proterebia'' Roos & Amschied, 1980 :* '' Loxerebia'' Watkins, 1925 :* '' Dyndirus'' Capronnier, 1874

Notes

References

Citations

Sources

* * * * * * * * ** * * **

External links

Satyrinae
at NYMPHALIDAE.net (There is no individual page for Satyrini as it is split into pages for each subtribes.)
Boisduval 1833
at TREE OF LIFE web project

at Pteron World (in Japanese) {{Taxonbar, from=Q2717036 Satyrinae Butterfly tribes