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Embolemidae is a family of small solitary parasitoid wasps with around 70 species in 2 genera distributed around the world.van Achterberg, Cornelis & Kats, R.. (2000). Revision of the Palaearctic Embolemidae (Hymenoptera). Zoöl. Med. 74 (2000), 17: 251-269. The few species whose biology is known are parasites on
planthopper A planthopper is any insect in the infraorder Fulgoromorpha, in the suborder Auchenorrhyncha, a group exceeding 12,500 described species worldwide. The name comes from their remarkable resemblance to leaves and other plants of their environment ...
nymphs of the families Achilidae and
Cixiidae The Cixiidae are a family of Fulgoroidea, fulgoroid insects, one of many families commonly known as planthoppers, distributed worldwide and comprising more than 2,000 species from over 150 Genus (biology), genera. Taxonomy Genera have been place ...
. There is debate regarding the status of the genus named ''Ampulicomorpha'' by Ashmead in 1893, generally considered now to be a junior synonym of '' Embolemus'' (e.g.,), though some authorities dispute this (e.g.,)


Biology

Females are wingless while males have wings, and in temperate regions emerge later than the females, which overwinter as adults. The wingless females have been recorded from the nests of ants and small mammal burrows, or under stones in pastures and grasslands, and they appear to be ant mimics. A Palearctic species, '' Embolemus ruddii'', has been found in association with the ant species '' Formica fusca'' and '' Lasius flavus'', while in Japan, '' Embolemus walkeri'' was taken in a nest of another ant, from the genus ''
Myrmica ''Myrmica'' is a genus of ants within the subfamily Myrmicinae. It is widespread throughout the temperate regions of the Holarctic realm, Holarctic and high mountains in Southeast Asia. The genus consists of around 200 known species and additio ...
''. A
Nearctic The Nearctic realm is one of the eight biogeographic realms constituting the Earth's land surface. The Nearctic realm covers most of North America, including Greenland, Central Florida, and the highlands of Mexico. The parts of North America ...
species, '' Embolemus confusus'', has been reared from nymphs of a
planthopper A planthopper is any insect in the infraorder Fulgoromorpha, in the suborder Auchenorrhyncha, a group exceeding 12,500 described species worldwide. The name comes from their remarkable resemblance to leaves and other plants of their environment ...
in the family Achilidae, where the host fed on fungi beneath the bark of rotting logs. The wasp larva lives in a bulging sac attached to the host nymph between the second and third segments.


Fossil species

After * †'' Baissobius'' Rasnitsyn 1975 ** †''Baissobius carolianus'' Rasnitsyn 1996 Dzun-Bain Formation, Mongolia, Early Cretaceous (
Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
) ** †''Baissobius minimus'' Rasnitsyn 1996
Zaza Formation The Zaza Formation is a geological formation located in Buryatia (Russia). It dates to the Lower Cretaceous. The age of the formation is disputed, and is considered likely to be Valanginian-Hauterivian, or Aptian in age. It comprises sandstones, ...
, Russia, Aptian ** †''Baissobius minutus'' Olmi et al. 2010 Zaza Formation, Russia, Aptian ** †''Baissobius parvus'' Rasnitsyn 1975 Zaza Formation, Russia, Aptian * †'' Cretembolemus'' Olmi et al. 2014 ** †''Cretembolemus orapensis'' Olmi et al. 2014
Orapa Orapa is a town located in the Central District of Botswana. It is the site of the Orapa diamond mine, the largest diamond-producing mine in the world, and is considered to be the diamond capital of the country. Nearby is another kimberlite ...
, Botswana, Late Cretaceous (
Turonian The Turonian is, in the International Commission on Stratigraphy, ICS' geologic timescale, the second age (geology), age in the Late Cretaceous epoch (geology), Epoch, or a stage (stratigraphy), stage in the Upper Cretaceous series (stratigraphy), ...
) * †'' Embolemopsis'' Olmi et al. 2010 ** †''Embolemopsis baissensis'' Olmi et al. 2010 Zaza Formation, Russia, Aptian ** †''Embolemopsis maryannae'' Olmi, Jarzembowski, Capradossi and Perkovsky, 2020
Wessex Formation The Wessex Formation is a fossil-rich England, English geological formation that dates from the Berriasian to Barremian Stage (stratigraphy), stages of the Early Cretaceous. It forms part of the Wealden Group and underlies the younger Vectis Form ...
, United Kingdom, Early Cretaceous (
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
) * '' Embolemus'' Westwood 1833 (incl. ''Ampulicomorpha'' Ashmead, 1893) ** †''Embolemus antiquus'' Perkovsky et al., 2021 Burmese amber, Myanmar, Late Cretaceous (
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
) ** †''Embolemus brachypterus'' Olmi et al., 2021 Burmese amber, Myanmar, Cenomanian ** †''Embolemus burmensis'' Perkovsky et al., 2021 Burmese amber, Myanmar, Cenomanian ** †''Embolemus cretacicus'' Perkovsky et al., 2021 Burmese amber, Myanmar, Cenomanian ** †''Embolemus excitus'' Perrichot and Engel 2011
Baltic amber Baltic amber or succinite is amber from the Baltic region, home of its largest known deposits. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that this forested region provided the re ...
,
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
** †''Embolemus janzeni'' (Olmi et al., 2014)
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. Th ...
, Myanmar, Cenomanian ** †''Embolemus micropterus'' Olmi et al., 2021 Burmese amber, Myanmar, Cenomanian ** †''Embolemus ohmkuhnlei'' Perkovsky et al., 2021 Burmese amber, Myanmar, Cenomanian ** †''Embolemus perialla'' (Ortega-Blanco et al., 2011) Spanish amber, Early Cretaceous (
Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
) ** †''Embolemus quesnoyensis'' (Chény, Guillam, Nel and Perrichot, 2020)
Oise amber Oise amber () is a type of amber found near the Oise river near Creil in northern France. Oise amber is around 53 million years old, dating to the Early Eocene (Ypresian). Oise amber is softer than Baltic amber, although Oise amber is older ...
, France, Eocene (
Ypresian In the geologic timescale the Ypresian is the oldest age (geology), age or lowest stage (stratigraphy), stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by th ...
) ** †''Embolemus succinalis'' (Brues, 1933) Baltic amber, Rovno amber, Eocene ** †''Embolemus zherikhini'' Perkovsky et al., 2021 Burmese amber, Myanmar, Cenomanian * †'' Ponomarenkoa'' Olmi 2010 ** †''Ponomarenkoa'' burmensis Perkovsky et al., 2021 Burmese amber, Myanmar, Cenomanian ** †''Ponomarenkoa ellenbergeri'' Olmi et al. 2013 Burmese amber, Myanmar, Cenomanian ** †''Ponomarenkoa polonica'' Ponomarenko 1988 Baltic amber, Eocene


References

{{Authority control Apocrita families Chrysidoidea