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''Changchunsaurus'' (meaning "
Changchun Changchun (, ; ), Chinese postal romanization, also romanized as Ch'angch'un, is the capital and largest city of Jilin, Jilin Province, China, People's Republic of China. Lying in the center of the Songliao Plain, Changchun is administered as a ...
lizard") is an extinct
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of small
herbivorous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23  million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
from
Early Cretaceous The Early Cretaceous (geochronological name) or the Lower Cretaceous ( chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Pr ...
deposits of
Gongzhuling Gongzhuling () is a county-level city under the administration of Changchun. It is located in central Jilin province of Northeast China, halfway between Siping and Changchun, along the main railway line in the Northeast. Major employers in the ci ...
,
Jilin Jilin (; alternately romanized as Kirin or Chilin) is one of the three provinces of Northeast China. Its capital and largest city is Changchun. Jilin borders North Korea ( Rasŏn, North Hamgyong, Ryanggang and Chagang) and Russia (P ...
, China. It is the first named dinosaur genus from Jilin.


Description

''Changchunsaurus'' was first named by Zan Shu-Qin, Chen Jun, Jin Li-Yong and Li Tao in
2005 File:2005 Events Collage V2.png, From top left, clockwise: Hurricane Katrina in the Gulf of Mexico; the Funeral of Pope John Paul II is held in Vatican City; "Me at the zoo", the first video ever to be uploaded to YouTube; Eris (dwarf planet), Er ...
. The
type Type may refer to: Science and technology Computing * Typing, producing text via a keyboard, typewriter, etc. * Data type, collection of values used for computations. * File type * TYPE (DOS command), a command to display contents of a file. * Ty ...
and only known
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
is ''C. parvus'' ("parvus" meaning "petite"), named for its small size. It is known from a skull and skeleton and additional skull fragments. All specimens of ''Changchunsaurus'' were collected from the Quantou Formation of the Songliao Basin, dating to the
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ag ...
Cenomanian stages. ''Changchunsaurus'' is based on the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
JLUM L0403-j–Zn2, a
skeleton A skeleton is the structural frame that supports the body of an animal. There are several types of skeletons, including the exoskeleton, which is the stable outer shell of an organism, the endoskeleton, which forms the support structure inside ...
and
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, t ...
, with a
premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has ...
(upper beak) and partial
lower jaw In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...
also known. Only the skull was figured and described in the official description. According to Zan ''et al.'' 2005, who described it, the animal shows a combination of features like those of
derived Derive may refer to: * Derive (computer algebra system), a commercial system made by Texas Instruments * ''Dérive'' (magazine), an Austrian science magazine on urbanism *Dérive, a psychogeographical concept See also * *Derivation (disambiguatio ...
ornithopods (reduction in size or loss of some skull
fenestrae A fenestra (fenestration; plural fenestrae or fenestrations) is any small opening or pore, commonly used as a term in the biology, biological sciences. It is the Latin word for "window", and is used in various fields to describe a pore in an anat ...
or holes), and features like those of more basal ornithopods (for example, five teeth in each premaxilla, short toothless portion of upper beak, and a small gap between beak teeth and cheek teeth). There is a projection that sticks out from the side of the
jugal The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by species. Anat ...
or cheekbone, with what is described as a "nubble structure". The type individual was a small animal, around 1 meter long (3.3 feet, with a skull 11.5 centimeters long (4.5 inches). It was originally classified as a basal ornithopod,
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
unknown Unknown or The Unknown may refer to: Film * ''The Unknown'' (1915 comedy film), a silent boxing film * ''The Unknown'' (1915 drama film) * ''The Unknown'' (1927 film), a silent horror film starring Lon Chaney * ''The Unknown'' (1936 film), a ...
, although it was not included in a formal
phylogenetic analysis In biology Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that pro ...
. Later, some referred specimens were described and in 2010 its
cranial Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...
anatomy was revised.


Classification

In Butler ''et al.'', 2011, the
postcrania Postcrania (postcranium, adjective: postcranial) in zoology and vertebrate paleontology is all or part of the skeleton apart from the skull. Frequently, fossil remains, e.g. of dinosaurs or other extinct tetrapods, consist of partial or isolated s ...
l osteology was described for the first time and a large
phylogenetic analysis In biology Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that pro ...
confirmed its position as a basal ornithopod which was found to be closely related to another Chinese ornithopod, '' Jeholosaurus'' and later to the newly described Chinese ornithopod ''
Haya Haya may refer to: Biology * ''Haya'' (dinosaur), a genus of basal ornithopod dinosaur that lived during the Late Cretaceous from Mongolia * ''Haya'' (plant), a genus of plants in the family Caryophyllaceae * Haya de Herguijuela (Spanish: beech ...
''. Han ''et al.'' named this clade "Jeholosauridae" in 2012. The following
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
was based on analysis by Makovicky ''et al.'', 2011.


References


External links


''Changchunsaurus''
at Dinosaur.net.cn, with the article summary and photograph. (slow-loading) {{Portal bar, Dinosaurs, Cretaceous, China Cretaceous dinosaurs of Asia Fossil taxa described in 2005 Ornithischian genera