Chaco Earthcreeper
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The Chaco earthcreeper (''Tarphonomus certhioides'') is a species of
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
in the Furnariinae subfamily of the ovenbird
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Furnariidae. It is found in
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
,
Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...
, and
Paraguay Paraguay, officially the Republic of Paraguay, is a landlocked country in South America. It is bordered by Argentina to the Argentina–Paraguay border, south and southwest, Brazil to the Brazil–Paraguay border, east and northeast, and Boli ...
, and as a
vagrant Vagrancy is the condition of wandering homelessness without regular employment or income. Vagrants usually live in poverty and support themselves by travelling while engaging in begging, scavenging, or petty theft. In Western countries, ...
in Brazil.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, G. Del-Rio, A. Jaramillo, D. F. Lane, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 31 May 2023. Species Lists of Birds for South American Countries and Territories. https://www.museum.lsu.edu/~Remsen/SACCCountryLists.htm retrieved May 31, 2023


Taxonomy and systematics

The Chaco earthcreeper has at times been placed in genera ''
Ochetorhynchus ''Ochetorhynchus'' is a genus of earthcreeper (disambiguation), earthcreepers, birds in the family Furnariidae. They are found in open to semi-open habitats in Chile, Argentina, Bolivia and Peru. The genus contains the following species: * Roc ...
'' and ''
Upucerthia ''Upucerthia'' is a genus of bird in the family Furnariidae. Taxonomy and etymology ''Upucerthia'' is a genus of bird in the family Furnariidae. Established by French zoologist Isidore Geoffroy Saint-Hilaire in 1832, it contains four species ...
'' but since the early 2000s has been placed in its current ''Tarphonomus''.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, G. Del-Rio, A. Jaramillo, D. F. Lane, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 31 May 2023. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved May 31, 2023 It has at times been considered
conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organism ...
with the other member of ''Tarphonomus'', the Bolivian earthcreeper (''T. harterti''). The two of them are
sisters A sister is a woman or a girl who shares parents or a parent with another individual; a female sibling. The male counterpart is a brother. Although the term typically refers to a familial relationship, it is sometimes used endearingly to r ...
to the rusty-winged barbtail (''Premnornis guttuliger''), and these three are in turn sisters to the tuftedcheeks of genus '' Pseudocolaptes''. The Chaco earthcreeper has three subspecies, the
nominate Nomination is part of the process of selecting a candidate for either election to a public office, or the bestowing of an honor or award. A collection of nominees narrowed from the full list of candidates is a short list. Political office In th ...
''T. c. certhoides'' (
d'Orbigny Alcide Charles Victor Marie Dessalines d'Orbigny (6 September 1802 – 30 June 1857) was a French naturalist who made major contributions in many areas, including zoology (including malacology), palaeontology, geology, archaeology and anthropol ...
& Lafresnaye, 1838), ''T. c. luscinia'' ( Burmeister, 1860), and ''T. c. estebani'' ( Wetmore & Peters, JL, 1949).


Description

The Chaco earthcreeper is about long and weighs . It is a small earthcreeper with a long and very slightly decurved bill. The sexes' plumages are alike. Adults of the nominate subspecies have an indistinct orange-rufous
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also k ...
on a mostly dark brown face. Their forehead is dull orange-rufous, their crown and back dull dark brown, and their rump and uppertail
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are small feathers behind t ...
somewhat
rufescent Rufous () is a color that may be described as reddish-brown or brownish-red, as of rust or oxidised iron. The first recorded use of ''rufous'' as a color name in English was in 1782. However, the color is also recorded earlier in 1527 as a dia ...
brown. Their tail is generally dull brown with progressively more rufous from the central to the outermost feathers. Their wings are slightly more rufous brown than the back with rufous bases on the flight feathers. Their throat and cheeks are white. Their breast and belly are brown and their flanks and undertail coverts rufescent. Their iris is brown, their
maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
blackish to dark gray, their
mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...
slate gray to pinkish gray, and their legs and feet blackish to dark gray. Juveniles have an overall rufous tinge but their forehead is less rufous than adults'.Remsen, Jr., J. V. (2020). Chaco Earthcreeper (''Tarphonomus certhioides''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.chaear1.01 retrieved August 20, 2023 Subspecies ''T. c. luscinia'' has grayer (less rufescent) upperparts, more rufous on the wings, and paler underparts than the nominate. ''T. c. estebani'' has a paler back, a duller breast and belly, and no rufescent tinge on the flanks compared to the nominate.


Distribution and habitat

Subspecies ''T. c. estebani'' of the Chaco earthcreeper is found in Santa Cruz Department in south central Bolivia, in northern Argentina, and in western Paraguay. ''T. c. luscinia'' is found in the western Argentina provinces of
San Juan San Juan, Spanish for Saint John (disambiguation), Saint John, most commonly refers to: * San Juan, Puerto Rico * San Juan, Argentina * San Juan, Metro Manila, a highly urbanized city in the Philippines San Juan may also refer to: Places Arge ...
,
La Rioja La Rioja () is an autonomous communities in Spain, autonomous community and provinces of Spain, province in Spain, in the north of the Iberian Peninsula. Its capital is Logroño. Other List of municipalities in La Rioja, cities and towns in the ...
,
Córdoba Córdoba most commonly refers to: * Córdoba, Spain, a major city in southern Spain and formerly the imperial capital of Islamic Spain * Córdoba, Argentina, the second largest city in Argentina and the capital of Córdoba Province Córdoba or Cord ...
, Mendoza, and San Luis. The nominate ''T. c. certhoides'' is found in northeastern Argentina from
Río Negro Province Río Negro (, ''Black River'') is a province of Argentina, located in northern Patagonia. Neighboring provinces are from the south clockwise Chubut, Neuquén, Mendoza, La Pampa and Buenos Aires. To the east lies the Atlantic Ocean. Its cap ...
north to
Formosa Province Formosa Province () is a province in northeastern Argentina, part of the Gran Chaco Region. Formosa's northeast end touches Asunción, Paraguay, and the province borders the provinces of Chaco and Salta to its south and west, respectively. The ...
. It has also occurred once in
Rio Grande do Sul Rio Grande do Sul (, ; ; "Great River of the South") is a Federative units of Brazil, state in the South Region, Brazil, southern region of Brazil. It is the Federative units of Brazil#List, fifth-most populous state and the List of Brazilian s ...
, Brazil.Repenning, M., Chiarani, E., Pereira, M.S. and Fontana, C.S. (2012). First record of the Chaco Earthcreeper ''Tarphonomus certhioides'' (Furnariidae) in Brazil. Rev. Bras. Orn. 20(4): 453–454. Subspecies ''T. c. estebani'' and ''T. c. certhoides'' of the Chaco earthcreeper inhabit dense scrublands and deciduous woodland in the
Chaco Basin The Chaco Basin (, or ) is a major sedimentary basin in Central South America around the borders of Argentina, Bolivia and Paraguay. The basin forms part of the larger Paraná Basin. Superficially, the Chaco Basin is an alluvial basin composed of ...
, especially areas with terrestrial
bromeliad The Bromeliaceae (the bromeliads) are a Family (biology), family of monocot flowering plants of about 80 genera and 3700 known species, native mainly to the Tropics, tropical Americas, with several species found in the American subtropics and on ...
s. ''T. c. luscinia'' inhabits slopes with dense shrubs in the Andean foothills. In elevation the species occurs up to .


Behavior


Movement

The Chaco earthcreeper is a year-round resident throughout its range.


Feeding

The Chaco earthcreeper feeds on
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s. It usually forages alone, gleaning prey from the ground and possibly from low vegetation.


Breeding

The Chaco earthcreeper breeds during the austral summer. It is thought to be monogamous. It builds a stick nest in a bush or tree, in a cavity in a tree, rock face, or old building, or in an old nest of a '' Furnarius'' hornero. It pads the floor of the nest with softer material such as hair, grass, and other plant fibers. The clutch size is three or four eggs. The incubation period, time to fledging, and details of parental care are not known.


Vocalization

The Chaco earthcreeper's song is "a series of 5–15 loud, squeaky penetrating 'chiqui' notes that increase in intensity nddescend in pitch". Its call is "a loud whistled 'tééoo' or 'tuéét' ".


Status

The IUCN has assessed the Chaco earthcreeper as being of Least Concern. It has a large range, and though its population size is not known it is believed to be stable. No immediate threats have been identified. It is considered fairly common in most of its range and occurs in at least one protected area.


References

{{Taxonbar, from=Q1274631 Chaco earthcreeper Birds of the Gran Chaco Chaco earthcreeper Taxonomy articles created by Polbot