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Steppe Pika
The steppe pika (''Ochotona pusilla'') is a small mammal of the pika family, Ochotonidae. It is found in the steppes of southern Russia and northern Kazakhstan. General description Ochotonidae includes only one genus - ''Ochotona'', formed by 30 living species. Like rodents, pikas have chisel-like incisor teeth, but they also have a second pair of incisors in the upper jaw, followed by two molar teeth in the upper jaw and three molar teeth in the lower jaw. Pikas have no canines. Their teeth grow throughout their life and they need to be worn down. The anatomical structure of the lower jaw bone makes it possible to move the jaw both up and down and sideways. Pikas have sharp senses, they can smell, hear and see very well. Their fibular bone has partially accreted with the tibia. They have five digits on their limbs and pads on feet and toes. Pikas are not hibernating animals. The steppe pika is usually 14.5-18.5 cm long. The tail is reduced, and the short round ears have ...
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Peter Simon Pallas
Peter Simon Pallas FRS FRSE (22 September 1741 – 8 September 1811) was a Prussian zoologist and botanist who worked in Russia between 1767 and 1810. Life and work Peter Simon Pallas was born in Berlin, the son of Professor of Surgery Simon Pallas. He studied with private tutors and took an interest in natural history, later attending the University of Halle and the University of Göttingen. In 1760, he moved to the University of Leiden and passed his doctor's degree at the age of 19. Pallas travelled throughout the Netherlands and to London, improving his medical and surgical knowledge. He then settled at The Hague, and his new system of animal classification was praised by Georges Cuvier. Pallas wrote ''Miscellanea Zoologica'' (1766), which included descriptions of several vertebrates new to science which he had discovered in the Dutch museum collections. A planned voyage to southern Africa and the East Indies fell through when his father recalled him to Berlin. ...
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Central Asia
Central Asia, also known as Middle Asia, is a region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes the former Soviet republics of Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, and Uzbekistan, which are colloquially referred to as the "-stans" as the countries all have names ending with the Persian suffix " -stan", meaning "land of". The current geographical location of Central Asia was formerly part of the historic region of Turkistan, also known as Turan. In the pre-Islamic and early Islamic eras ( and earlier) Central Asia was inhabited predominantly by Iranian peoples, populated by Eastern Iranian-speaking Bactrians, Sogdians, Chorasmians and the semi-nomadic Scythians and Dahae. After expansion by Turkic peoples, Central Asia also became the homeland for the Kazakhs, Uzbeks, Tatars, Turkmen, Kyrgyz, and Uyghurs; ...
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Owl Pellet
A pellet, in ornithology, is the mass of undigested parts of a bird's food that some bird species occasionally regurgitate. The contents of a bird's pellet depend on its diet, but can include the exoskeletons of insects, indigestible plant matter, bones, fur, feathers, bills, claws, and teeth. In falconry, the pellet is called a ''casting''. The passing of pellets allows a bird to remove indigestible material from its proventriculus, or glandular stomach. In birds of prey, the regurgitation of pellets serves the bird's health in another way, by "scouring" parts of the digestive tract, including the gullet. Pellets are formed within six to ten hours of a meal in the bird's gizzard (muscular stomach). Ornithologists may collect one species' pellets over time to analyze the seasonal variation in its eating habits. One advantage of collecting pellets is that it allows for the determination of diet without the killing and dissection of the bird. Pellets are found in different loca ...
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Fossil Record
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the absol ...
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Holocene
The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene together form the Quaternary period. The Holocene has been identified with the current warm period, known as MIS 1. It is considered by some to be an interglacial period within the Pleistocene Epoch, called the Flandrian interglacial.Oxford University Press – Why Geography Matters: More Than Ever (book) – "Holocene Humanity" section https://books.google.com/books?id=7P0_sWIcBNsC The Holocene corresponds with the rapid proliferation, growth and impacts of the human species worldwide, including all of its written history, technological revolutions, development of major civilizations, and overall significant transition towards urban living in the present. The human impact on modern-era Earth and its ecosystems may be considered of gl ...
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Chalcolithic Europe
The European Chalcolithic, the Chalcolithic (also Eneolithic, Copper Age) period of Prehistoric Europe, lasted roughly from 5000 to 2000 BC, developing from the preceding Neolithic period. It was a period of Megalithic culture, the appearance of the first significant economic stratification, and probably the earliest presence of Indo-European speakers. The economy of the Chalcolithic, even in the regions where copper was not yet used, was no longer that of peasant communities and tribes: some materials began to be produced in specific locations and distributed to wide regions. Mining of metal and stone was particularly developed in some areas, along with the processing of those materials into valuable goods. Ancient Chalcolithic From c. 5000 BC to 3000 BC, copper started being used in the Balkans, Eastern Europe, and Central Europe. From c. 3500 onwards, there was an influx of people into Eastern Europe from the area east of the Volga (Yamnaya culture), creating a plural co ...
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Carpathian Basin
The Pannonian Basin, or Carpathian Basin, is a large Sedimentary basin, basin situated in south-east Central Europe. The Geomorphology, geomorphological term Pannonian Plain is more widely used for roughly the same region though with a somewhat different sense, with only the Upland and lowland, lowlands, the plain that remained when the Pliocene Epoch ''Pannonian Sea'' dried out. It is a geomorphological subsystem of the Alpide belt, Alps-Himalaya system, specifically a sediment-filled back-arc basin which divergent boundary, spread apart during the Miocene. The plain or basin is diagonally bisected by the Transdanubian Mountains, separating the larger Great Hungarian Plain (including the Eastern Slovak Lowland) from the Little Hungarian Plain. It forms a topographically discrete unit set in the European landscape, surrounded by imposing geographic boundaries—the Carpathian Mountains and the Alps. The Rivers Danube and Tisza divide the basin roughly in half. It extends rough ...
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Pleistocene
The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in 2009 by the International Union of Geological Sciences, the cutoff of the Pleistocene and the preceding Pliocene was regarded as being 1.806 million years Before Present (BP). Publications from earlier years may use either definition of the period. The end of the Pleistocene corresponds with the end of the last glacial period and also with the end of the Paleolithic age used in archaeology. The name is a combination of Ancient Greek grc, label=none, πλεῖστος, pleīstos, most and grc, label=none, καινός, kainós (latinized as ), 'new'. At the end of the preceding Pliocene, the previously isolated North and South American continents were joined by the Isthmus of Panama, causing Great American Interchang ...
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Urals
The Ural Mountains ( ; rus, Ура́льские го́ры, r=Uralskiye gory, p=ʊˈralʲskʲɪjə ˈɡorɨ; ba, Урал тауҙары) or simply the Urals, are a mountain range that runs approximately from north to south through western Russia, from the coast of the Arctic Ocean to the river Ural and northwestern Kazakhstan.Ural Mountains
Encyclopædia Britannica on-line
The mountain range forms part of the conventional boundary between the regions of and

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MtDNA
Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA is only a small portion of the DNA in a eukaryotic cell; most of the DNA can be found in the cell nucleus and, in plants and algae, also in plastids such as chloroplasts. Human mitochondrial DNA was the first significant part of the human genome to be sequenced. This sequencing revealed that the human mtDNA includes 16,569 base pairs and encodes 13 proteins. Since animal mtDNA evolves faster than nuclear genetic markers, it represents a mainstay of phylogenetics and evolutionary biology. It also permits an examination of the relatedness of populations, and so has become important in anthropology and biogeography. Origin Nuclear and mitochondrial DNA are thought to be of separate evolutionary origin, with the mtDNA being derived from t ...
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Mammal
Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or hair, and three middle ear bones. These characteristics distinguish them from reptiles (including birds) from which they diverged in the Carboniferous, over 300 million years ago. Around 6,400 extant species of mammals have been described divided into 29 orders. The largest orders, in terms of number of species, are the rodents, bats, and Eulipotyphla ( hedgehogs, moles, shrews, and others). The next three are the Primates (including humans, apes, monkeys, and others), the Artiodactyla (cetaceans and even-toed ungulates), and the Carnivora ( cats, dogs, seals, and others). In terms of cladistics, which reflects evolutionary history, mammals are the only living members of the Synapsida (synapsids); this clade, toget ...
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Karyology
A karyotype is the general appearance of the complete set of metaphase chromosomes in the cells of a species or in an individual organism, mainly including their sizes, numbers, and shapes. Karyotyping is the process by which a karyotype is discerned by determining the chromosome complement of an individual, including the number of chromosomes and any abnormalities. A karyogram or idiogram is a graphical depiction of a karyotype, wherein chromosomes are organized in pairs, ordered by size and position of centromere for chromosomes of the same size. Karyotyping generally combines light microscopy and photography, and results in a photomicrographic (or simply micrographic) karyogram. In contrast, a schematic karyogram is a designed graphic representation of a karyotype. In schematic karyograms, just one of the sister chromatids of each chromosome is generally shown for brevity, and in reality they are generally so close together that they look as one on photomicrographs as well u ...
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