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Eupelycosaur
Eupelycosauria is a large clade of animals characterized by the unique shape of their skull, encompassing all mammals and their closest extinct relatives. They first appeared 308million years ago during the Early Pennsylvanian epoch, with the fossils of ''Echinerpeton'' and perhaps an even earlier genus, ''Protoclepsydrops'', representing just one of the many stages in the evolution of mammals,Kemp. T.S., 1982, ''Mammal-like Reptiles and the Origin of Mammals''. Academic Press, New York in contrast to their earlier amniote ancestors. Taxonomy Eupelycosaurs are synapsids, animals whose skull has a single opening behind the eye. They are distinguished from the Caseasaurian synapsids by having a long, narrow supratemporal bone (instead of one that is as wide as it is long) and a frontal bone with a wider connection to the upper margin of the orbit. Laurin, M. and Reisz, R. R., 1997Autapomorphies of the main clades of synapsids- Tree of Life Web Project The only living descendant ...
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Synapsid
Synapsida is a diverse group of tetrapod vertebrates that includes all mammals and their extinct relatives. It is one of the two major clades of the group Amniota, the other being the more diverse group Sauropsida (which includes all extant reptiles and therefore, birds). Unlike other amniotes, synapsids have a single temporal fenestra, an opening low in the skull roof behind each eye socket, leaving a zygomatic arch, bony arch beneath each; this accounts for the name "synapsid". The distinctive temporal fenestra developed about 318 million years ago during the Late Carboniferous period, when synapsids and sauropsids diverged, but was subsequently merged with the orbit in early mammals. The basal (phylogenetics), basal amniotes (reptiliomorphs) from which synapsids evolved were historically simply called "reptiles". Therefore, stem group synapsids were then described as mammal-like reptiles in classical systematics, and non-therapsid synapsids were also referred to as pelyco ...
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Mammal
A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three Evolution of mammalian auditory ossicles, middle ear bones. These characteristics distinguish them from reptiles and birds, from which their ancestors Genetic divergence, diverged in the Carboniferous Period over 300 million years ago. Around 6,640 Neontology#Extant taxon, extant species of mammals have been described and divided into 27 Order (biology), orders. The study of mammals is called mammalogy. The largest orders of mammals, by number of species, are the rodents, bats, and eulipotyphlans (including hedgehogs, Mole (animal), moles and shrews). The next three are the primates (including humans, monkeys and lemurs), the Artiodactyl, even-toed ungulates (including pigs, camels, and whales), and the Carnivora (including Felidae, ...
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Pennsylvanian (geology)
The Pennsylvanian ( , also known as Upper Carboniferous or Late Carboniferous) is, on the International Commission on Stratigraphy, ICS geologic timescale, the younger of two period (geology), subperiods of the Carboniferous Period (or the upper of two system (stratigraphy), subsystems of the Carboniferous System). It lasted from roughly . As with most other geochronology, geochronologic units, the stratum, rock beds that define the Pennsylvanian are well identified, but the exact date of the start and end are uncertain by a few hundred thousand years. The Pennsylvanian is named after the U.S. state of Pennsylvania, where the coal Bed (geology), beds of this age are widespread. The division between Pennsylvanian and Mississippian (geology), Mississippian comes from North American stratigraphy. In North America, where the early Carboniferous beds are primarily marine limestones, the Pennsylvanian was in the past treated as a full-fledged geologic period between the Mississippian ...
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Echinerpeton
''Echinerpeton'' is an extinct genus of synapsid, including the single species ''Echinerpeton intermedium'' from the Late Carboniferous of Nova Scotia, Canada. The name means 'spiny lizard' (Greek). Along with its contemporary ''Archaeothyris'', ''Echinerpeton'' is the oldest known synapsid, having lived around 308 million years ago. It is known from six small, fragmentary fossils, which were found in an outcrop of the Morien Group near the town of Florence. The most complete specimen preserves articulated vertebrae with high neural spines, indicating that ''Echinerpeton'' was a sail-backed synapsid like the better known ''Dimetrodon'', ''Sphenacodon'', and ''Edaphosaurus''. However, the relationship of ''Echinerpeton'' to these other forms is unclear, and its phylogenetic placement among basal synapsids remains uncertain. Description ''Echinerpeton'' is known from six specimens, five housed in the Museum of Comparative Zoology and a sixth in the Redpath Museum: the holotype MCZ ...
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Tree Of Life Web Project
The Tree of Life Web Project (ToL) is an Internet project providing information about the diversity and phylogeny of life on Earth. This collaborative peer reviewed project began in 1995, and is written by biologists from around the world. The site has not been updated since 2011, however the pages are still accessible. The pages are linked hierarchically, in the form of the branching evolutionary tree of life, organized cladistically. Each page contains information about one particular group of organisms and is organized according to a branched tree-like form, thus showing hypothetical relationships between different groups of organisms. In 2009 the project ran into funding problems from the University of Arizona. Pages and Treehouses submitted took a considerably longer time to be approved as they were being reviewed by a small group of volunteers, and apparently, around 2011, all activities ended. History The idea of this project started in the late 1980s. David Maddison ...
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Robert Reisz
Robert Rafael Reisz is a Canadian paleontologist and specialist in the study of early amniote and tetrapod evolution. Research career Reisz received his B.Sc. (1969), M.Sc. (1971) and Ph.D. (1975) from McGill University as Robert L. Carroll's first doctoral graduate. After teaching as visiting lecturer at University of California, Los Angeles for a year, he accepted an appointment in the Biology Department at the University of Toronto's Mississauga Campus in 1975 where he still maintains his research lab. His research has been funded continuously by the Natural Sciences and Engineering Research Council of Canada (NSERC). He conducted field work in North America, Africa, and Europe, where he excavated fossils from the late Paleozoic and early Mesozoic eras. These excavations were frequently funded by the National Geographic Society. Reisz has broad interests in vertebrate paleontology. He has published more than 100 scientific articles on subjects as diverse as lungfish and di ...
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Michel Laurin
Michel Laurin is a Canadian-born French vertebrate paleontologist whose specialities include the emergence of a land-based lifestyle among vertebrates, the evolution of body size and the origin and phylogeny of lissamphibians. He has also made important contributions to the literature on phylogenetic nomenclature. As an undergraduate, he worked in the laboratory of Robert L. Carroll and earned his Ph.D. at the University of Toronto under the direction of Robert R. Reisz; his thesis concerned the osteology of Seymouriamorpha, seymouriamorphs. His 1991 review of diapsid phylogeny provided the broadest review of the subject up to that date. In 1995, Laurin and Reisz coauthored a widely cited article providing evidence that the synapsids are the sister group of all other amniotes. He later worked on untangling the phylogeny of the Stegocephalia, a group with a notoriously difficult phylogeny. He later moved to France; since 1998, he has been a CNRS researcher at the Muséum National d'H ...
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Orbit (anatomy)
In anatomy Anatomy () is the branch of morphology concerned with the study of the internal structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old scien ..., the orbit is the Body cavity, cavity or socket/hole of the skull in which the eye and Accessory visual structures, its appendages are situated. "Orbit" can refer to the bony socket, or it can also be used to imply the contents. In the adult human, the volume of the orbit is about , of which the eye occupies . The orbital contents comprise the eye, the Orbital fascia, orbital and retrobulbar fascia, extraocular muscles, cranial nerves optic nerve, II, oculomotor nerve, III, trochlear nerve, IV, trigeminal nerve, V, and abducens nerve, VI, blood vessels, fat, the lacrimal gland with its Lacrimal sac, sac and nasolacrimal duct, duct, the eyelids, Medial palpebral ligament, medial and Lateral palpebral raphe, lateral palpebr ...
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Frontal Bone
In the human skull, the frontal bone or sincipital bone is an unpaired bone which consists of two portions.'' Gray's Anatomy'' (1918) These are the vertically oriented squamous part, and the horizontally oriented orbital part, making up the bony part of the forehead, part of the bony orbital cavity holding the eye, and part of the bony part of the nose respectively. The name comes from the Latin word ''frons'' (meaning "forehead"). Structure The frontal bone is made up of two main parts. These are the squamous part, and the orbital part. The squamous part marks the vertical, flat, and also the biggest part, and the main region of the forehead. The orbital part is the horizontal and second biggest region of the frontal bone. It enters into the formation of the roofs of the orbital and nasal cavities. Sometimes a third part is included as the nasal part of the frontal bone, and sometimes this is included with the squamous part. The nasal part is between the brow ridges, ...
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Supratemporal Bone
The supratemporal bone is a paired cranial bone present in many tetrapods and tetrapodomorph fish. It is part of the temporal region (the portion of the skull roof behind the eyes), usually lying medial (inwards) relative to the squamosal and lateral (outwards) relative to the parietal and/or postparietal. It may also contact the postorbital or intertemporal (which lie forwards), or tabular (which lies backwards), when those bones are present. The supratemporal is a common component of the skull in many extinct amphibians, though it is apparently absent in the lightweight skulls of living lissamphibians (frogs and salamanders). Embryological studies of salamanders suggests that the supratemporal fuses with the squamosal in early development. A separate supratemporal was retained by early synapsids and reptiles, but was strongly reduced in many groups. Squamates (lizards and snakes) still possess a small supratemporal, though archosaurs (crocodilians and birds) and mammals lack i ...
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Caseasauria
Caseasauria is one of the two main clades of early synapsids, the other being the Eupelycosauria. Caseasaurs are currently known only from the Late Carboniferous and the Permian, and include two superficially different families, the small insectivorous or carnivorous Eothyrididae, and the large, herbivory, herbivorous Caseidae. These two groups share a number of specialised features associated with the morphology of the snout and external nares, naris. The ancestor of caseasaurs can be traced back to an insect eating or an omnivorous reptile-like synapsid from the Pennsylvanian (geology), Pennsylvanian time of the Carboniferous, possibly resembling ''Archaeothyris'', the earliest known synapsid. The caseasaurs were abundant during the later part of the Early Permian, but by the Middle Permian caseasaur diversity declined because the group was outcompeted by the more successful therapsids. The last caseasaurs became extinct at the end of the Guadelupian (Middle Permian). Descript ...
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Synapomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel Phenotypic trait, character or character state that has evolution, evolved from its ancestral form (or Plesiomorphy and symplesiomorphy, plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxon, taxa and is therefore Hypothesis#Scientific hypothesis, hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies Homology (biology), homology. Examples of apomorphy are the presence of Terrestrial locomotion#Posture, erect gait, fur, Evolution of mammalian auditory ossicles, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a Terrestrial locomotion#Posture, sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word ...
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