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Chasmosaurines
Chasmosaurinae is a subfamily of ceratopsid dinosaurs. They were one of the most successful groups of herbivores of their time. Chasmosaurines appeared in the early Campanian, and became extinct, along with all other non-avian dinosaurs, during the Cretaceous–Paleogene extinction event. Broadly, the most distinguishing features of chasmosaurines are prominent brow horns and long frills lacking long spines; centrosaurines generally had short brow horns and relatively shorter frills, and often had long spines projecting from their frills. Chasmosaurines evolved in western North America (Laramidia). They are currently known definitively from rocks in western Canada, the western United States, and northern Mexico. They were highly diverse and among the most species-rich groups of dinosaurs, with new species frequently described. This high diversity of named species is likely a result of the frill. The distinctive shape of the frill with the hornlets on its edges (epoccipitals) make ...
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Kosmoceratops
''Kosmoceratops'' () is a genus of ceratopsid dinosaur that lived in North America about 76–75.9 million years ago during the Late Cretaceous period. Specimens were discovered in Utah in the Kaiparowits Formation of the Grand Staircase–Escalante National Monument in 2006 and 2007, including an adult skull and postcranial skeleton and partial subadults. In 2010, the adult was made the holotype of the new genus and species ''Kosmoceratops richardsoni''; the generic name means "ornate horned face", and the specific name honors Scott Richardson, who found the specimens. The find was part of a spate of ceratopsian discoveries in the early 21st century, and ''Kosmoceratops'' was considered significant due to its elaborate skull ornamentation. ''Kosmoceratops'' had an estimated length of and a weight of . As a ceratopsid, it would have been quadrupedal with a heavily constructed skeleton. It had a triangular beak with a pointed tip and a blade-like nasal horn with a flattened u ...
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Bravoceratops
''Bravoceratops'' is a genus of large chasmosaurine ceratopsid dinosaur that lived approximately 70 million years ago, and is known from the Late Cretaceous Javelina Formation in what is now Texas, United States. Discovery and naming ''Bravoceratops'' is only known from the holotype specimen TMM 46015-1; it is housed in the collection of the Texas Memorial Museum, Austin, Texas. The skull consists of a number of fragments, altogether including: the , parts of each brow horn, the rear end of the left , assorted parts of the nasal area and horn, the , and quadratojugals from each side, a section of the and , multiple parts of the dentary, and some of the right . It was recovered from the lowermost rocks of the Javelina Formation, in Big Bend National Park; ceratopsid fossils are uncommon in this formation, giving the discovery of ''Bravoceratops'' importance in clarifying the groups' diversity. The specimen was found in sandy conglomerate sediment at the Hippiewalk geologic loc ...
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Anchiceratops
''Anchiceratops'' ( ) is an extinct genus of chasmosaurinae, chasmosaurine ceratopsid dinosaur that lived approximately 72 to 71 million years ago during the latter part of the Cretaceous Period (geology), Period in what is now Alberta, Canada. ''Anchiceratops'' was a medium-sized, heavily built, ground-dwelling, quadrupedal herbivore that could grow up to an estimated long. Its skull featured two long brow horns and a short horn on the nose. The skull frill was elongated and rectangular, its edges adorned by coarse triangular projections. About a dozen skulls of the genus have been found. History of discovery The first remains of ''Anchiceratops'' were discovered along the Red Deer River in the Provinces and territories of Canada, Canadian province of Alberta in 1912 by an expedition led by Barnum Brown. The holotype, specimen AMNH 5251, is the back half of a skull, including the long frill, and two other partial skulls, specimens AMNH 5259 (the paratype) and AMNH 5273, were f ...
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Agujaceratops
''Agujaceratops'' (meaning "horned face from Aguja") is a genus of horned dinosaur from the Late Cretaceous (Campanian) of west Texas. It is a chasmosaurine (long-frilled) ceratopsian. Two species are known, ''Agujaceratops mariscalensis'', and ''A''. ''mavericus''. Discovery and species In 1938, three dinosaur bone beds were excavated, and ceratopsian material was collected from Big Bend National Park (Texas) by William Strain. This material was studied by Lehman in 1989 and named ''Chasmosaurus mariscalensis''. It is known only from the holotype UTEP P.37.7.086 a partial adult skull which includes a braincase, left supraorbital horncore, left maxilla and a right dentary. Additional material was associated with the holotype, but not considered to be part of it. All specimens of ''Agujaceratops'' were collected from the lower part of the Upper Shale member of the Aguja Formation, dating to about 77 million years ago, in the Big Bend National Park, Brewster County. Additio ...
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Titanoceratops
''Titanoceratops'' (meaning "titanic horned face") is a controversial genus of herbivorous ceratopsian dinosaur. It was a giant chasmosaurine ceratopsian that lived in the Late Cretaceous period (Campanian stage, about 75 million years agoFowler, D. W. 2017. Revised geochronology, correlation, and dinosaur stratigraphic ranges of the Santonian-Maastrichtian (Late Cretaceous) formations of the Western Interior of North America. PLoS ONE 12(11): e0188426.) in what is now New Mexico. ''Titanoceratops'' was named for its large size, being one of the largest known horned dinosaurs and the type species was named ''T. ouranos'', after Uranus (Ouranos), the father of the Greek titans. It was named in 2011 by Nicholas R. Longrich for a specimen previously referred to '' Pentaceratops''. Longrich believed that unique features found in the skull reveal it to have been a close relative of ''Triceratops'', classified within the subgroup Triceratopsini. However, other researchers have expres ...
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Utahceratops
''Utahceratops'' is an extinct genus of ceratopsian dinosaur that lived approximately 76.4~75.5 million years ago during the Late Cretaceous period in what is now Utah. ''Utahceratops'' was a large-sized, robustly-built, ground-dwelling, quadrupedal herbivore, that could grow up to an estimated long. Discovery The genus name ''Utahceratops'', means "horned face from Utah", and is derived from the state of Utah and Greek words "keras" (κέρας) meaning "horn" and "ops" (ὤψ) referring to the "face". The specific name ''gettyi'', is derived from the name of Mike Getty, who discovered the holotype and has played a pivotal role in the recovery of fossils from the Grand Staircase–Escalante National Monument (GSENM). It was first named by Scott D. Sampson, Mark A. Loewen, Andrew A. Farke, Eric M. Roberts, Catherine A. Forster, Joshua A. Smith and Alan L. Titus in 2010, and the type species is ''Utahceratops gettyi''. Description The holotype specimen UMNH VP 16784, consist ...
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Chasmosaurus Belli
''Chasmosaurus'' ( ) is a genus of ceratopsid dinosaur from the Upper Cretaceous Period of North America. Its name means 'opening lizard', referring to the large openings (fenestrae) in its frill (Greek ''chasma'' meaning 'opening' or 'hollow' or 'gulf' and ''sauros'' meaning 'lizard'). With a length of and a weight of , ''Chasmosaurus'' was a ceratopsian of average size. Like all ceratopsians, it was purely herbivorous. It was initially to be called ''Protorosaurus'', but this name had been previously published for another animal. All specimens of ''Chasmosaurus'' were collected from the Dinosaur Park Formation of the Dinosaur Provincial Park of Alberta, Canada. Referred specimens of ''C. russelli'' come from the lower beds of the formation while ''C. belli'' comes from middle and upper beds. Discovery and species In 1898, at ''Berry Creek'', Alberta, Lawrence Morris Lambe of the Geological Survey of Canada made the first discovery of ''Chasmosaurus'' remains; holotype NMC 4 ...
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Dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23  million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is the subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya; their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic, morphological and ecological standpoints. Birds, at over 10,700 living species ...
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Ceratopsidae
Ceratopsidae (sometimes spelled Ceratopidae) is a family of ceratopsian dinosaurs including ''Triceratops'', ''Centrosaurus'', and ''Styracosaurus''. All known species were quadrupedal herbivores from the Upper Cretaceous. All but one species are known from western North America, which formed the island continent of Laramidia during most of the Late Cretaceous. Ceratopsids are characterized by beaks, rows of shearing teeth in the back of the jaw, elaborate nasal horns, and a thin parietal-squamosal shelf that extends back and up into a frill. The group is divided into two subfamilies— Chasmosaurinae and Centrosaurinae. The chasmosaurines are generally characterized by long, triangular frills and well-developed brow horns. The centrosaurines had well-developed nasal horns or nasal bosses, shorter and more rectangular frills, and elaborate spines on the back of the frill. These horns and frills show remarkable variation and are the principal means by which the various species have ...
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Vagaceratops
''Vagaceratops'' (meaning "wandering (''vagus'', Latin) horned face", in reference to its close relationship with '' Kosmoceratops'' from Utah) is a genus of herbivorous ceratopsian dinosaur. It is a chasmosaurine ceratopsian which lived during the Late Cretaceous period (late Campanian) in what is now Alberta. Its fossils have been recovered from the Upper Dinosaur Park Formation. It is sometimes included in the genus ''Chasmosaurus'' as ''Chasmosaurus irvinensis'' instead of being recognized as its own genus. Description ''Vagaceratops'' was a medium-sized ceratopsian, reaching in length and weighing . It is known primarily from three fossil skulls. Although the general structure was typical of ceratopsids (i.e. a parrot-like beak, large neck frill, and nasal horn) it has some peculiarities. The skulls are characterized by a reduced supraorbital horn, brow horns that are reduced to low bosses and a larger snout compared to related animals. ''Vagaceratops'' had smaller pari ...
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Spiclypeus
''Spiclypeus'' (meaning "spike shield") is an extinct genus of chasmosaurine ceratopsian dinosaur known from the Late Cretaceous Judith River Formation (late Campanian stage) of Montana, United States. Discovery In 2000, Bill D. Shipp, a nuclear physicist, bought the Paradise Point Ranch near the town of Winifred, in Fergus County. Believing that his land contained fossils, Shipp hired the local veteran fossil collector John C. Gilpatrick to explore the terrain together. On their first trip during an afternoon in September 2005, Shipp found the ''Spiclypeus'' specimen on his land in Montana. He saw a thighbone jutting out of a hillside at the Judith River Breaks. Shipp then hired the amateur paleontologist Joe Small to excavate the fossils. At the cost of several hundred thousand dollars, a road was constructed to allow an excavator to remove the overburden covering the skull of the specimen. In 2007, Small and his team managed to secure all remaining bones. The fossils were pr ...
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Chasmosaurus
''Chasmosaurus'' ( ) is a genus of ceratopsid dinosaur from the Upper Cretaceous Period of North America. Its name means 'opening lizard', referring to the large openings (fenestrae) in its frill (Greek ''chasma'' meaning 'opening' or 'hollow' or 'gulf' and ''sauros'' meaning 'lizard'). With a length of and a weight of , ''Chasmosaurus'' was a ceratopsian of average size. Like all ceratopsians, it was purely herbivorous. It was initially to be called ''Protorosaurus'', but this name had been previously published for another animal. All specimens of ''Chasmosaurus'' were collected from the Dinosaur Park Formation of the Dinosaur Provincial Park of Alberta, Canada. Referred specimens of ''C. russelli'' come from the lower beds of the formation while ''C. belli'' comes from middle and upper beds. Discovery and species In 1898, at ''Berry Creek'', Alberta, Lawrence Morris Lambe of the Geological Survey of Canada made the first discovery of ''Chasmosaurus'' remains; holotype NM ...
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