''Agujaceratops'' (meaning "horned face from Aguja") is a
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of horned
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
from the Late Cretaceous (
Campanian
The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campani ...
) of west
Texas
Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
. It is a
chasmosaurine (long-frilled)
ceratopsian
Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassi ...
. Two species are known, ''Agujaceratops mariscalensis'', and ''A''. ''mavericus''.
Discovery and species

In 1938, three dinosaur bone beds were excavated, and ceratopsian material was collected from
Big Bend National Park
Big Bend National Park is an American national park located in West Texas, bordering Mexico. The park has national significance as the largest protected area of Chihuahuan Desert topography and ecology in the United States, and was named after ...
(Texas) by William Strain. This material was studied by Lehman in 1989 and named ''Chasmosaurus mariscalensis''. It is known only from the
holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
UTEP
The University of Texas at El Paso (UTEP) is a public research university in El Paso, Texas. It is a member of the University of Texas System. UTEP is the second-largest university in the United States to have a majority Mexican American stude ...
P.37.7.086 a partial adult
skull
The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, t ...
which includes a
braincase
In human anatomy, the neurocranium, also known as the braincase, brainpan, or brain-pan is the upper and back part of the skull, which forms a protective case around the brain. In the human skull, the neurocranium includes the calvaria or skul ...
, left supraorbital
horncore
A horn is a permanent pointed projection on the head of various animals that consists of a covering of keratin and other proteins surrounding a core of live bone. Horns are distinct from antlers, which are not permanent.
In mammals, true horns ...
, left
maxilla
The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The ...
and a right
dentary
In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...
. Additional material was associated with the holotype, but not considered to be part of it.
All specimens of ''Agujaceratops'' were collected from the lower part of the Upper Shale member of the
Aguja Formation
The Aguja Formation is a geological formation in North America, exposed in Texas, United States and Chihuahua and Coahuila in Mexico, whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered f ...
, dating to about 77
million years ago
The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds.
Usage
Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...
,
in the
Big Bend National Park
Big Bend National Park is an American national park located in West Texas, bordering Mexico. The park has national significance as the largest protected area of Chihuahuan Desert topography and ecology in the United States, and was named after ...
,
Brewster County
Brewster County is a county located in the U.S. state of Texas. It is in West Texas and its county seat (and only city) is Alpine. It is one of the nine counties that comprise the Trans-Pecos region, and borders Mexico. Brewster County is t ...
. Additional material was recovered from elsewhere in west Texas, including a nearly complete skull from Rattlesnake Creek.
Originally described as ''
Chasmosaurus
''Chasmosaurus'' ( ) is a genus of ceratopsid dinosaur from the Upper Cretaceous Period of North America. Its name means 'opening lizard', referring to the large openings (fenestrae) in its frill (Greek ''chasma'' meaning 'opening' or 'hollow' ...
mariscalensis'' by Lehman in
1989
File:1989 Events Collage.png, From left, clockwise: The Cypress structure collapses as a result of the 1989 Loma Prieta earthquake, killing motorists below; The proposal document for the World Wide Web is submitted; The Exxon Valdez oil tanker ru ...
, subsequent analysis resulted in the taxon being put in its own genus. ''Agujaceratops'' was named by Spencer G. Lucas, Robert M. Sullivan and Adrian Hunt in
2006, and the
type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
is ''Agujaceratops mariscalensis''.
Later, Lehman and colleagues revisited the ''Agujaceratops'' material and found substantial variation. They described the Rattlesnake Creek skull as a new species, ''Agujaceratops mavericus''.
Description

''Agujaceratops'' was a relatively large horned dinosaur, reaching in length and in body mass.
It was similar to other chasmosaurines such as ''
Pentaceratops
''Pentaceratops'' ("five-horned face") is a genus of herbivorous ceratopsid dinosaur from the late Cretaceous Period of what is now North America. Fossils of this animal were first discovered in 1921, but the genus was named in 1923 when its t ...
'' in having a short nose horn, long brow horns, and an elongate frill circled by small hornlets. The back of the frill has a strong notch, as in ''
Pentaceratops
''Pentaceratops'' ("five-horned face") is a genus of herbivorous ceratopsid dinosaur from the late Cretaceous Period of what is now North America. Fossils of this animal were first discovered in 1921, but the genus was named in 1923 when its t ...
'' and ''
Chasmosaurus
''Chasmosaurus'' ( ) is a genus of ceratopsid dinosaur from the Upper Cretaceous Period of North America. Its name means 'opening lizard', referring to the large openings (fenestrae) in its frill (Greek ''chasma'' meaning 'opening' or 'hollow' ...
'', giving it a heart shape, with three or four pairs of spike-like hornlets. The edges of the frill bear numerous low, blunt hornlets, giving it a strongly scalloped appearance. The brow horns are oriented up and out, and curve backwards in side view.

Two species are known, ''Agujaceratops mariscalensis'' and ''A''. ''mavericus''. ''A''. ''mariscalensis'' has shorter brow horns and a shorter frill.
Ecology
Like other ceratopsids, ''Agujaceratops'' was a four-legged plant eater. The elaborate frill and horns suggest a complex social life, perhaps involving displays towards and fights with other members of the species over territory or mating. Multiple individuals are found in a single quarry. It is unclear whether this represents animals brought together by a drought or flood event, or perhaps a herd. Although it is common to find multiple individuals of
centrosaurine
Centrosaurinae (from the Greek, meaning "pointed lizards") is a subfamily of ceratopsid dinosaurs, a group of large quadrupedal ornithischians. Centrosaurine fossil remains are known primarily from the northern region of Laramidia (modern day ...
ceratopsids together - large bonebeds are known for ''
Centrosaurus
''Centrosaurus'' ( ; ) is a genus of herbivorous ceratopsian dinosaur from the Late Cretaceous of Canada. Their remains have been found in the Dinosaur Park Formation, dating from 76.5 to 75.5 million years ago.
Discovery and naming
Th ...
'' and ''
Pachyrhinosaurus
''Pachyrhinosaurus'' (meaning in Greek "thick-nosed lizard", from ' (), thick; ' (), nose; and (), lizard) is an extinct genus of centrosaurine ceratopsid dinosaur from the Late Cretaceous period of North America. The first examples were disco ...
'', for example - bonebeds are rarer for chasmosaurines.
At the time, the Aguja Formation lay along the western margin of the
Western Interior Seaway
The Western Interior Seaway (also called the Cretaceous Seaway, the Niobraran Sea, the North American Inland Sea, and the Western Interior Sea) was a large inland sea that split the continent of North America into two landmasses. The ancient se ...
. The habitat ''Agujaceratops'' lived in (at least where the fossil material was found) may have been a swamp, due to the nature of the sediments.
''Agujaceratops'' lived alongside a fauna that included the
feathered dinosaur
A feathered dinosaur is any species of dinosaur possessing feathers. While this includes all species of birds, there is a hypothesis that many, if not all non-avian dinosaur species also possessed feathers in some shape or form.
It has been s ...
''
Leptorhynchos gaddisi'' and the small
pachycephalosaur ''
Texacephale
''Texacephale'' is a pachycephalosaurid dinosaur from the Campanian stage of the Late Cretaceous. Its fossils come from the Aguja Formation of Big Bend National Park, in Texas, and were described in 2010 by Longrich, Sankey and Tanke. The generi ...
''. Predators would have included
tyrannosaurs
Tyrannosauroidea (meaning 'tyrant lizard forms') is a superfamily (or clade) of coelurosaurian theropod dinosaurs that includes the family Tyrannosauridae as well as more basal relatives. Tyrannosauroids lived on the Laurasian supercontinent ...
and the giant
crocodilia
Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest livi ...
n ''
Deinosuchus
''Deinosuchus'' () is an extinct genus of alligatoroid crocodilian, related to modern alligators and caimans, that lived 82 to 73 million years ago (Ma), during the late Cretaceous period. The name translates as "terrible crocodile" and ...
riograndensis''.
See also
*
Timeline of ceratopsian research
This timeline of ceratopsian research is a chronological listing of events in the History of paleontology, history of paleontology focused on the ceratopsians, a group of herbivorous marginocephalian dinosaurs that evolved parrot-like beaks, bon ...
*
2016 in paleontology
Flora Plants
Fungi
Cnidarians
Research
* '' Yunnanoascus haikouensis'', previously thought to be a member of Ctenophora, is reinterpreted as a crown-group medusozoan by Han ''et al.'' (2016).
* A study on the fossil corals from the Late T ...
References
*
External links
DinoGeorge's List of Dinosaurs, including ''Agujaceratops'' an
{{Taxonbar, from=Q132534
Chasmosaurines
Late Cretaceous dinosaurs of North America
Fossil taxa described in 2006
Taxa named by Spencer G. Lucas
Paleontology in Texas
Campanian genus first appearances
Campanian genus extinctions
Ornithischian genera