Tropeognathus Mesembrinus
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Tropeognathus Mesembrinus
''Tropeognathus'' (meaning "keel jaw") is a genus of large pterosaurs from the late Early Cretaceous of South America. This genus is considered to be a member of the family Anhangueridae, however, several studies have also recovered it within another family called Ornithocheiridae. Both of these families are diverse groups of pterosaurs known for their keel-tipped snouts and large size. ''Tropeognathus'' is regarded as the largest pterosaur found in the Southern Hemisphere, only rivaled by the huge azhdarchids. The type and only species is ''Tropeognathus mesembrinus''. Fossil remains of ''Tropeognathus'' have been recovered from the Romualdo Formation, which is a Lagerstätte located in the Santana Group of the Araripe Basin in northeastern Brazil. Discovery and naming In the 1980s the German paleontology museum '' Bayerische Staatssammlung für Paläontologie und historische Geologie'' in Munich acquired a pterosaur skull from Brazilian fossil dealers that had probably b ...
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Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145 Megaannum#SI prefix multipliers, Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian-Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining Anoxic event#Cretaceous, early Aptian Oceanic Anoxic Event 1a (OAE1a) was carbon isotope dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java Plateau, Ontong Java-Manihiki Plateau, Manihiki-Hikurangi Plateau, Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. T ...
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Lagerstätte
A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These formations may have resulted from carcass burial in an anoxic environment with minimal bacteria, thus delaying the decomposition of both gross and fine biological features until long after a durable impression was created in the surrounding matrix. ''Lagerstätten'' span geological time from the Neoproterozoic era to the present. Worldwide, some of the best examples of near-perfect fossilization are the Cambrian Maotianshan shales and Burgess Shale, the Silurian Waukesha Biota, the Devonian Hunsrück Slates and Gogo Formation, the Carboniferous Mazon Creek, the Jurassic Posidonia Shale and Solnhofen Limestone, the Cretaceous Yixian, Santana, and Agua Nueva formations, the Eocene Green River Formation, the Miocene Foulden Maar ...
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Holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering. A holotype is not necessarily " ...
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Koine
Koine Greek (; Koine el, ἡ κοινὴ διάλεκτος, hē koinè diálektos, the common dialect; ), also known as Hellenistic Greek, common Attic, the Alexandrian dialect, Biblical Greek or New Testament Greek, was the common supra-regional form of Greek spoken and written during the Hellenistic period, the Roman Empire and the early Byzantine Empire. It evolved from the spread of Greek following the conquests of Alexander the Great in the fourth century BC, and served as the lingua franca of much of the Mediterranean region and the Middle East during the following centuries. It was based mainly on Attic and related Ionic speech forms, with various admixtures brought about through dialect levelling with other varieties. Koine Greek included styles ranging from conservative literary forms to the spoken vernaculars of the time. As the dominant language of the Byzantine Empire, it developed further into Medieval Greek, which then turned into Modern Greek. Literary Koi ...
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Specific Name (zoology)
In zoological nomenclature, the specific name (also specific epithet or species epithet) is the second part (the second name) within the scientific name of a species (a binomen). The first part of the name of a species is the name of the genus or the generic name. The rules and regulations governing the giving of a new species name are explained in the article species description. For example, the scientific name for humans is ''Homo sapiens'', which is the species name, consisting of two names: ''Homo'' is the " generic name" (the name of the genus) and ''sapiens'' is the "specific name". Historically, ''specific name'' referred to the combination of what are now called the generic and specific names. Carl Linnaeus, who formalized binomial nomenclature, made explicit distinctions between specific, generic, and trivial names. The generic name was that of the genus, the first in the binomial, the trivial name was the second name in the binomial, and the specific the proper term fo ...
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Type Species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen(s). Article 67.1 A similar concept is used for suprageneric groups and called a type genus. In botanical nomenclature, these terms have no formal standing under the code of nomenclature, but are sometimes borrowed from zoological nomenclature. In botany, the type of a genus name is a specimen (or, rarely, an illustration) which is also the type of a species name. The species name that has that type can also be referred to as the type of the genus name. Names of genus and family ranks, the various subdivisions of those ranks, and some higher-rank names based on genus names, have such types.
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Chapada Do Araripe
The Chapada do Araripe, also known as the Serra do Araripe, is a ''chapada'' ( plateau) in northeastern Brazil. The chapada forms the boundary of Ceará and Pernambuco states, and forms the watershed between the Jaguaribe River of Ceará, which flows northward into the Atlantic, and the much larger basin of the São Francisco River of Pernambuco and neighboring states, which drains eastward into the Atlantic. The Santana Formation, which is rich with fossils from the early Cretaceous, lies at the base of the chapada. The Araripe manakin The Araripe manakin (''Antilophia bokermanni'') is a species of critically endangered bird from the family of manakins (Pipridae). It was discovered in 1996 and scientifically described in 1998. The species epithet commemorates Brazilian zoologi ... (''Antilophia bokermanni''), a bird only described in 1998, is endemic to Chapada do Araripe. Etymology The word "Araripe" stems from the Tupi word ''ararype'', meaning " macaw at the river" (' ...
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Ceará
Ceará (, pronounced locally as or ) is one of the 26 states of Brazil, located in the northeastern part of the country, on the Atlantic coast. It is the eighth-largest Brazilian State by population and the 17th by area. It is also one of the main tourist destinations in Brazil. The state capital is the city of Fortaleza, the country's fourth most populous city. The state has 4.3% of the Brazilian population and produces 2.1% of the Brazilian GDP. Literally, the name ''Ceará'' means "sings the jandaia". According to José de Alencar, one of the most important writers of Brazil and an authority in Tupi Guaraní, ''Ceará'' means turquoise or green waters. The state is best known for its extensive coastline, with of sand. There are also mountains and valleys producing tropical fruits. To the south, on the border of Paraíba, Pernambuco and Piauí, is the National Forest of Araripe. Geography Ceará has an area of . It is bounded on the north by the Atlantic Ocean ...
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Munich
Munich ( ; german: München ; bar, Minga ) is the capital and most populous city of the German state of Bavaria. With a population of 1,558,395 inhabitants as of 31 July 2020, it is the third-largest city in Germany, after Berlin and Hamburg, and thus the largest which does not constitute its own state, as well as the 11th-largest city in the European Union. The city's metropolitan region is home to 6 million people. Straddling the banks of the River Isar (a tributary of the Danube) north of the Bavarian Alps, Munich is the seat of the Bavarian administrative region of Upper Bavaria, while being the most densely populated municipality in Germany (4,500 people per km2). Munich is the second-largest city in the Bavarian dialect area, after the Austrian capital of Vienna. The city was first mentioned in 1158. Catholic Munich strongly resisted the Reformation and was a political point of divergence during the resulting Thirty Years' War, but remained physically unt ...
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Paleontology
Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossils to classify organisms and study their interactions with each other and their environments (their paleoecology). Paleontological observations have been documented as far back as the 5th century BC. The science became established in the 18th century as a result of Georges Cuvier's work on comparative anatomy, and developed rapidly in the 19th century. The term itself originates from Greek (, "old, ancient"), (, (gen. ), "being, creature"), and (, "speech, thought, study"). Paleontology lies on the border between biology and geology, but differs from archaeology in that it excludes the study of anatomically modern humans. It now uses techniques drawn from a wide range of sciences, including biochemistry, mathematics, and engineeri ...
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Tropeognathus
''Tropeognathus'' (meaning "keel jaw") is a genus of large pterosaurs from the late Early Cretaceous of South America. This genus is considered to be a member of the family Anhangueridae, however, several studies have also recovered it within another family called Ornithocheiridae. Both of these families are diverse groups of pterosaurs known for their keel-tipped snouts and large size. ''Tropeognathus'' is regarded as the largest pterosaur found in the Southern Hemisphere, only rivaled by the huge azhdarchids. The type and only species is ''Tropeognathus mesembrinus''. Fossil remains of ''Tropeognathus'' have been recovered from the Romualdo Formation, which is a Lagerstätte located in the Santana Group of the Araripe Basin in northeastern Brazil. Discovery and naming In the 1980s the German paleontology museum '' Bayerische Staatssammlung für Paläontologie und historische Geologie'' in Munich acquired a pterosaur skull from Brazilian fossil dealers that had probably be ...
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