Lagerstätte
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A Lagerstätte (, from ''
Lager Lager () is beer which has been brewed and conditioned at low temperature. Lagers can be pale, amber, or dark. Pale lager is the most widely consumed and commercially available style of beer. The term "lager" comes from the German for "storag ...
'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a
sedimentary Sedimentary rocks are types of rock (geology), rock that are formed by the accumulation or deposition of mineral or organic matter, organic particles at Earth#Surface, Earth's surface, followed by cementation (geology), cementation. Sedimentati ...
deposit that exhibits extraordinary
fossils A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ...
with exceptional preservation—sometimes including preserved soft tissues. These formations may have resulted from carcass burial in an
anoxic The term anoxia means a total depletion in the level of oxygen, an extreme form of hypoxia or "low oxygen". The terms anoxia and hypoxia are used in various contexts: * Anoxic waters, sea water, fresh water or groundwater that are depleted of diss ...
environment with minimal bacteria, thus delaying the decomposition of both gross and fine biological features until long after a durable impression was created in the surrounding matrix. ''Lagerstätten'' span
geological Geology () is a branch of natural science concerned with Earth and other astronomical objects, the features or rocks of which it is composed, and the processes by which they change over time. Modern geology significantly overlaps all other E ...
time from the
Neoproterozoic The Neoproterozoic Era is the unit of geologic time from 1 billion to 538.8 million years ago. It is the last era of the Precambrian Supereon and the Proterozoic Eon; it is subdivided into the Tonian, Cryogenian, and Ediacaran periods. It is ...
era to the
present The present (or here'' and ''now) is the time that is associated with the events perception, perceived directly and in the first time, not as a recollection (perceived more than once) or a speculation (predicted, hypothesis, uncertain). It is ...
. Worldwide, some of the best examples of near-perfect fossilization are the
Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...
Maotianshan shales The Maotianshan Shales are a series of Cambrian, Early Cambrian deposits in the Chiungchussu Formation, famous for their ''Lagerstätte, Konservat Lagerstätten'', deposits known for the exceptional preservation of fossilized organisms or traces. ...
and
Burgess Shale The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At old (middle Cambrian), it is one of the earliest foss ...
, the
Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozo ...
Waukesha Biota The Waukesha Biota (also known as Waukesha Lagerstätte, Brandon Bridge Lagerstätte, or Brandon Bridge fauna) refers to a Konservat-Lagerstätte ( a fossil site that preserves soft bodied remains) of Early Silurian ( Telychian to Sheinwoodian) ...
, the
Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ...
Hunsrück Slates and
Gogo Formation The Gogo Formation in the Kimberley region of Western Australia is a Lagerstätte that exhibits exceptional preservation of a Devonian reef community. The formation is named after Gogo Station, a cattle station where outcrops appear and fossil ...
, the
Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...
Mazon Creek The Mazon Creek fossil beds are a conservation ' found near Morris, in Grundy County, Illinois. The fossils are preserved in ironstone concretions, formed approximately in the mid- Pennsylvanian epoch of the Carboniferous period. These concreti ...
, the
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
Posidonia Shale The Posidonia Shale (german: Posidonienschiefer, also called Schistes Bitumineux in Luxembourg) geologically known as the Sachrang Formation, is an Early Jurassic (Toarcian) geological formation of southwestern and northeast Germany, northern Swit ...
and Solnhofen Limestone, the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
Yixian,
Santana Santana may refer to: Transportation * Volkswagen Santana, an automobile * Santana Cycles, manufacturer of tandem bicycles * Santana Motors, a former Spanish automobile manufacturer Boats * Santana 20, an American sailboat design by W. D. Sch ...
, and Agua Nueva formations, the
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
Green River Formation The Green River Formation is an Eocene geologic formation that records the sedimentation in a group of intermountain lakes in three basins along the present-day Green River in Colorado, Wyoming, and Utah. The sediments are deposited in very fine ...
, the
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
Foulden Maar Foulden Maar is a fossil site near Middlemarch in Otago, New Zealand. The fossils were deposited in the small deep crater lake of a maar formed by a volcano in the Miocene era, around 23 million years ago. The crater lake existed for a period of ...
and
Ashfall Fossil Beds The Ashfall Fossil Beds of Antelope County in northeastern Nebraska are rare fossil sites of the type called lagerstätten that, due to extraordinary local conditions, capture an ecological "snapshot" in time of a range of well-preserved fossil ...
, the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Gray Fossil Site The Gray Fossil Site is an Early Pliocene assemblage of fossils dating between 4.5 and 4.9 million years old, located near the town of Gray in Washington County, Tennessee. The site was discovered during a widening project of Tennessee State Route ...
, the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
Naracoorte Caves Naracoorte Caves National Park is a national park near Naracoorte in the Limestone Coast tourism region in the south-east of South Australia ( Australia). It was officially recognised in 1994 for its extensive fossil record when the site was in ...
, the La Brea Tar Pits, and the Tanis Fossil Site.


Types

Palaeontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
s distinguish two kinds:The term was originally coined by
Adolf Seilacher Adolf "Dolf" Seilacher (February 24, 1925 – April 26, 2014) was a German palaeontologist who worked in evolutionary and ecological palaeobiology for over 60 years. He is best known for his contributions to the study of trace fossils; constructi ...
in:
# ''Konzentrat-Lagerstätten'' (concentration ''Lagerstätten'') are deposits with a particular "concentration" of disarticulated organic hard parts, such as a bone bed. These ''Lagerstätten'' are less spectacular than the more famous ''Konservat-Lagerstätten''. Their contents invariably display a large degree of time averaging, as the accumulation of bones in the absence of other sediment takes some time. Deposits with a high concentration of fossils that represent an ''in situ'' community, such as
reef A reef is a ridge or shoal of rock, coral or similar relatively stable material, lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic processes— deposition of sand, wave erosion planing down rock out ...
s or
oyster Oyster is the common name for a number of different families of salt-water bivalve molluscs that live in marine or brackish habitats. In some species, the valves are highly calcified, and many are somewhat irregular in shape. Many, but not ...
beds, are not considered ''Lagerstätten''. # ''Konservat-Lagerstätten'' (conservation ''Lagerstätten'') are deposits known for the exceptional preservation of fossilized organisms or traces. The individual
taphonomy Taphonomy is the study of how organisms decay and become fossilized or preserved in the paleontological record. The term ''taphonomy'' (from Greek , 'burial' and , 'law') was introduced to paleontology in 1940 by Soviet scientist Ivan Efremov t ...
of the fossils varies with the sites. Conservation ''Lagerstätten'' are crucial in providing answers to important moments in the history and
evolution Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
of life. For example, the
Burgess Shale The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At old (middle Cambrian), it is one of the earliest foss ...
of British Columbia is associated with the
Cambrian explosion The Cambrian explosion, Cambrian radiation, Cambrian diversification, or the Biological Big Bang refers to an interval of time approximately in the Cambrian Period when practically all major animal phyla started appearing in the fossil recor ...
, and the Solnhofen limestone with the earliest known
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
, '' Archaeopteryx''.


Preservation

''Konservat-Lagerstätten'' preserve lightly sclerotized and soft-bodied organisms or traces of organisms that are not otherwise preserved in the usual shelly and bony fossil record; thus, they offer more complete records of ancient biodiversity and behavior and enable some reconstruction of the palaeoecology of ancient aquatic communities. In 1986,
Simon Conway Morris Simon Conway Morris (born 1951) is an English palaeontologist, evolutionary biologist, and astrobiologist known for his study of the fossils of the Burgess Shale and the Cambrian explosion. The results of these discoveries were celebrated in ...
calculated only about 14% of genera in the Burgess Shale had possessed biomineralized tissues in life. The affinities of the shelly elements of
conodont Conodonts (Greek ''kōnos'', "cone", + ''odont'', "tooth") are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, which ...
s were mysterious until the associated soft tissues were discovered near Edinburgh, Scotland, in the Granton Lower
Oil Shale Oil shale is an organic-rich fine-grained sedimentary rock containing kerogen (a solid mixture of organic chemical compounds) from which liquid hydrocarbons can be produced. In addition to kerogen, general composition of oil shales constitute ...
of the
Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...
.Briggs et al. 1983; Aldridge et al. 1993. Information from the broader range of organisms found in ''Lagerstätten'' have contributed to recent
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
reconstructions of some major
metazoan Animals are multicellular, eukaryotic organisms in the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, and go through an ontogenetic stage in ...
groups. ''Lagerstätten'' seem to be temporally autocorrelated, perhaps because global environmental factors such as climate might affect their deposition. A number of
taphonomic Taphonomy is the study of how organisms decay and become fossilized or preserved in the paleontological record. The term ''taphonomy'' (from Greek , 'burial' and , 'law') was introduced to paleontology in 1940 by Soviet scientist Ivan Efremov t ...
pathways may produce ''Lagerstätten''. The following is an incomplete list: * Orsten-type and Doushantuo-type preservations preserve organisms in
phosphate In chemistry, a phosphate is an anion, salt, functional group or ester derived from a phosphoric acid. It most commonly means orthophosphate, a derivative of orthophosphoric acid . The phosphate or orthophosphate ion is derived from phospho ...
. * Bitter Springs-type preservation preserves them in
silica Silicon dioxide, also known as silica, is an oxide of silicon with the chemical formula , most commonly found in nature as quartz and in various living organisms. In many parts of the world, silica is the major constituent of sand. Silica is one ...
. * Carbonaceous films are the result of Burgess Shale-type preservation * Pyrite preserves exquisite detail in Beecher's trilobite-type preservation. * Ediacaran-type preservation preserves casts and moulds with the aid of
microbial mat A microbial mat is a multi-layered sheet of microorganisms, mainly bacteria and archaea, or bacteria alone. Microbial mats grow at interfaces between different types of material, mostly on submerged or moist surfaces, but a few survive in deserts ...
s.


Important ''Konservat-Lagerstätten''

The world's major ''Lagerstätten'' include:


See also

*
List of fossil sites This list of fossil sites is a worldwide list of localities known well for the presence of fossils. Some entries in this list are notable for a single, unique find, while others are notable for the large number of fossils found there. Many of t ...
(with link directory) * Hoard, a concentration of human artifacts useful for similar reasons in archaeology


References


Further reading

* Penney, D. (ed.) 2010. ''Biodiversity of Fossils in Amber from the Major World Deposits''. Siri Scienfic Press, Manchester, 304 pp. * – A catalogue of sites of exceptional fossil preservation produced by MSc palaeobiology students at University of Bristol's Department of Earth Sciences. * {{DEFAULTSORT:Lagerstatte German words and phrases