Lagerstätte

A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These formations may have resulted from carcass burial in an anoxic environment with minimal bacteria, thus delaying the decomposition of both gross and fine biological features until long after a durable impression was created in the surrounding matrix. ''Lagerstätten'' span geological time from the Neoproterozoic era to the present. Worldwide, some of the best examples of near-perfect fossilization are the Cambrian Maotianshan shales and Burgess Shale, the Silurian Waukesha Biota, the Devonian Hunsrück Slates and Gogo Formation, the Carboniferous Mazon Creek, the Jurassic Posidonia Shale and Solnhofen Limestone, the Cretaceous Yixian, Santana, and Agua Nueva formations, the Eocene Green River Formation, the Miocene Foulden Maar and Ashfall Fossil Beds, the Pliocene, the Pleistocene Naracoorte Caves, the La Brea Tar Pits, and the Tanis Fossil Site.

Types

Palaeontologists distinguish two kinds:The term was originally coined by Adolf Seilacher in:
# ''Konzentrat-Lagerstätten'' (concentration ''Lagerstätten'') are deposits with a particular "concentration" of disarticulated organic hard parts, such as a bone bed. These ''Lagerstätten'' are less spectacular than the more famous ''Konservat-Lagerstätten''. Their contents invariably display a large degree of time averaging, as the accumulation of bones in the absence of other sediment takes some time. Deposits with a high concentration of fossils that represent an ''in situ'' community, such as reefs or oyster beds, are not considered ''Lagerstätten''.
# ''Konservat-Lagerstätten'' (conservation ''Lagerstätten'') are deposits known for the exceptional preservation of fossilized organisms or traces. The individual taphonomy of the fossils varies with the sites. Conservation ''Lagerstätten'' are crucial in providing answers to important moments in the history and evolution of life. For example, the Burgess Shale of British Columbia is associated with the Cambrian explosion, and the Solnhofen limestone with the earliest known bird, '' Archaeopteryx''.

Preservation

''Konservat-Lagerstätten'' preserve lightly sclerotized and soft-bodied organisms or traces of organisms that are not otherwise preserved in the usual shelly and bony fossil record; thus, they offer more complete records of ancient biodiversity and behavior and enable some reconstruction of the palaeoecology of ancient aquatic communities. In 1986, Simon Conway Morris calculated only about 14% of genera in the Burgess Shale had possessed biomineralized tissues in life. The affinities of the shelly elements of conodonts were mysterious until the associated soft tissues were discovered near Edinburgh, Scotland, in the Granton Lower Oil Shale of the Carboniferous.Briggs et al. 1983; Aldridge et al. 1993. Information from the broader range of organisms found in ''Lagerstätten'' have contributed to recent phylogenetic reconstructions of some major metazoan groups. ''Lagerstätten'' seem to be temporally autocorrelated, perhaps because global environmental factors such as climate might affect their deposition.

A number of taphonomic pathways may produce ''Lagerstätten''. The following is an incomplete list:
* Orsten-type and Doushantuo-type preservations preserve organisms in phosphate.
* Bitter Springs-type preservation preserves them in silica.
* Carbonaceous films are the result of Burgess Shale-type preservation
* Pyrite preserves exquisite detail in Beecher's trilobite-type preservation.
* Ediacaran-type preservation preserves casts and moulds with the aid of microbial mats.

Important ''Konservat-Lagerstätten''

The world's major ''Lagerstätten'' include:

See also

* List of fossil sites (with link directory)
* Hoard, a concentration of human artifacts useful for similar reasons in archaeology

References

Further reading

* Penney, D. (ed.) 2010. ''Biodiversity of Fossils in Amber from the Major World Deposits''. Siri Scienfic Press, Manchester, 304 pp.
* – A catalogue of sites of exceptional fossil preservation produced by MSc palaeobiology students at University of Bristol's Department of Earth Sciences.
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German words and phrases