Titanosauromorpha
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Titanosauromorpha
Somphospondyli is an extinct clade of titanosauriform sauropods that lived from the Late Jurassic until the end of the Late Cretaceous, comprising all titanosauriforms more closely related to Titanosauria proper than Brachiosauridae. The remains of somphospondylans have been discovered in all six continents. Classification The group has officially been defined under the PhyloCode as the largest clade containing ''Saltasaurus loricatus'', but not ''Giraffatitan brancai.'' Features found as diagnostic of this clade by Mannion ''et al.'' (2013) include the possession of at least 15 cervical vertebrae; a bevelled radius bone end; sacral vertebrae with camellate internal texture; convex posterior articular surfaces of middle to posterior caudal vertebrae; biconvex distal caudal vertebrae; humerus anterolateral corner "squared"; among multiple others. The following cladogram depicts the reference phylogeny used to defined Somphospondlyi under the PhyloCode The ''Internationa ...
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Late Jurassic
The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name "Malm" indicates rocks of Late Jurassic age. In the past, ''Malm'' was also used to indicate the unit of geological time, but this usage is now discouraged to make a clear distinction between lithostratigraphic and geochronologic/chronostratigraphic units. Subdivisions The Late Jurassic is divided into three ages, which correspond with the three (faunal) stages of Upper Jurassic rock: Paleogeography During the Late Jurassic Epoch, Pangaea broke up into two supercontinents, Laurasia to the north, and Gondwana to the south. The result of this break-up was the spawning of the Atlantic Ocean. However, at this time, the Atlantic Ocean was relatively narrow. Life forms of the epoch This epoch is well known for many famous types of dinosau ...
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Gobititan
''Gobititan'' is a genus of herbivorous sauropod dinosaur from the Aptian faunal stage of the Early Cretaceous. The name of this genus is derived from the Gobi desert region and the Titans of Greek mythology, which is a reference to its large body size. The specific name ''shenzhouensis'', is derived from "Shenzhou", an ancient name for China. Description ''Gobititan'' can be distinguished from other titanosauriforms based on features of the caudal vertebrae. Compared with advanced titanosaurs, where the number of caudal vertebrae had been reduced to less than 35, ''Gobititan'' had a relatively high number of caudal vertebrae, which was interpreted as a basal trait. Gregory S. Paul estimated that ''Gobititan'' was long and weighed . Discovery and naming The genus is based on one partial skeleton, holotype IVPP 12579, which consists of a series of 41 caudal vertebrae and an incomplete left hindlimb. Its remains were recovered in the summer of 1999 at the "Middle Gray unit" ...
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Extinct
Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and recover may have been lost before this point. Because a species' potential range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxa, where a species presumed extinct abruptly "reappears" (typically in the fossil record) after a period of apparent absence. More than 99% of all species that ever lived on Earth, amounting to over five billion species, are estimated to have died out. It is estimated that there are currently around 8.7 million species of eukaryote globally, and possibly many times more if microorganisms, like bacteria, are included. Notable extinct animal species include non-avian dinosaurs, saber-toothed cats, dodos, ...
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Titanosauria
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as '' Patagotitan''—estimated at long with a weight of —and the comparably-sized '' Argentinosaurus'' and ''Puertasaurus'' from the same region. The group's name alludes to the mythological Titans of ancient Greek mythology, via the type genus (now considered a ''nomen dubium)'' ''Titanosaurus''. Together with the brachiosaurids and relatives, titanosaurs make up the larger sauropod clade Titanosauriformes. Titanosaurs have long been a poorly-known group, and the relationships between titanosaur species are still not well-understood. Description Titanosauria have the largest ...
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Euhelopodidae
Euhelopodidae is a family of sauropod dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...s of disputed membership and affinities, which contains '' Euhelopus'' and its close relatives. Most proposed euhelopodids are from East Asia. Euhelopodidae was first recognized by Carl Wiman in 1929, under the name Helopodidae, as ''Euhelopus'' was originally named ''Helopus''. However, the name had already been proposed for a bird, so in 1956 Alfred Sherwood Romer proposed the name ''Euhelopus'' and Euhelopodinae as replacements; Romer classified Euhelopodinae as a subfamily of Brachiosauridae, in which he also included Camarasaurinae and Cetiosaurinae, rather than as a family of its own. In addition to ''Euhelopus'' itself, Romer included ''Chiayusaurus'', ''Omeisaurus'', and ''Tie ...
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Wintonotitan
''Wintonotitan'' (meaning " Winton titan") is a genus of titanosauriform dinosaur from late Albian (Early Cretaceous)-age Winton Formation of Australia. It is known from partial postcranial remains. Description and history Fossils that are now known under the name ''Wintonotitan'' were first found in 1974 by Keith Watts. At the time, the specimens were assigned to an '' Austrosaurus'' sp., ''Austrosaurus'' then being the only named Australian Cretaceous sauropod genus. These fossils, catalogued as QMF 7292, consisted of a left shoulder blade, much of the forelimbs, a number of back, hip, and tail vertebrae, part of the right hip, ribs, chevrons, and unidentifiable fragments. QMF 7292 was established as the type specimen of ''Wintonotitan'' in 2009 by Scott Hocknull and colleagues. Hocknull suggested that ''Austrosaurus mckillopi'' differed only slightly from the QMF 7292, the holotype of ''Wintonotitan wattsii'', and should be considered a ''nomen dubium''. The t ...
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Triunfosaurus
''Triunfosaurus'' (meaning "Triunfo Basin reptile") is a genus of somphospondylan sauropod dinosaur from the Early Cretaceous of Brazil. It contains a single species, ''T. leonardii'', described by Carvalho ''et al.'' in 2017. As a genus, ''Triunfosaurus'' can be distinguished from all other titanosaurs by the unique proportions of its ischium. It was initially described as a basal titanosaur, making it the earliest basal titanosaur known; however, subsequent research questioned the identification of the taxon as a titanosaur, instead reassigning it to the Somphospondyli. Description ''Triunfosaurus'' can be distinguished from other titanosaurs by two autapomorphies, or unique traits that distinguish it from other titanosaurs. Namely, the part of the ischium touching the pubis is half the height of the entire ischium, and the shaft of the ischium makes an angle of less than 70° with the acetabular portion, which contributes to the hip socket. It also presents a unique combi ...
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Tastavinsaurus
''Tastavinsaurus'' is a genus of sauropod dinosaur belonging to the Titanosauriformes. It is based on a partial skeleton from the Early Cretaceous Xert Formation of Spain. The type species is ''Tastavinsaurus sanzi'', named in honor of the Rio Tastavins in Spain and Spanish paleontologist José Luis Sanz. ''Tastavinsaurus sanzi'' is known from two specimens, one from the Xert Formation and one from the Forcall Formation. ''Tastavinsaurus'' was originally described as a somphospondylan, but it may be a brachiosaurid. The clade Laurasiformes was coined for ''Tastavinsaurus'' and its close relatives, which may include ''Cedarosaurus'', '' Soriatitan'', and ''Venenosaurus ''Venenosaurus'' ( ) was a sauropod dinosaur. The name literally means "poison lizard", and it was named so after the Poison Strip Member of the Cedar Mountain Formation in Utah, United States, where the fossils were discovered by a Denver Museum ...''. References Macronarians Early Cretaceous dinosa ...
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Sibirotitan
''Sibirotitan'' ("Siberian titan") is a genus of somphospondyl sauropod from the Ilek Formation of Russia. The type and only species is ''S. astrosacralis''. Discovery and naming The material assigned to ''Sibirotitan'' was found in the Shestakovo 1 locality of the Ilek Formation, deposited in a cliff on the Kiya River's right bank, near Shestakovo Village in Kemerovo Province, West Siberia, Russia. Vertebrate remains were first recovered in 1953, and larger dinosaur fossils were found later in the sixties. Definitive sauropod remains were first discovered during expeditions in 1994 and 1995. In 2002, a reasonably complete foot would be described by Russian vertebrate paleontologist Alexander Averianov and colleagues; there was insufficient material to name the taxon, but they identified it as a member of Titanosauriformes, noting teeth from the locality indicated possible brachiosaur identity, but that a caudal vertebra from a nearby locality indicated the presence of a ...
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Sauroposeidon
''Sauroposeidon'' ( ; meaning "lizard earthquake god", after the Greek god Poseidon) is a genus of sauropod dinosaur known from several incomplete specimens including a bone bed and fossilized trackways that have been found in the U.S. states of Oklahoma, Wyoming, and Texas. The fossils were found in rocks dating from near the end of the Early Cretaceous (Aptian–early Albian), from about 113 to 110 million years ago, a time when sauropod diversity in North America had greatly diminished. It was the last known North American sauropod prior to an absence of the group on the continent of roughly 40 million years that ended with the appearance of ''Alamosaurus'' during the Maastrichtian. While the holotype remains were initially discovered in 1994, due to their unexpected age and unusual size they were initially misclassified as pieces of petrified wood. A more detailed analysis in 1999 revealed their true nature which resulted in a minor media frenzy, and formal publication of th ...
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Ruixinia
''Ruixinia'' is an extinct genus of somphospondylan titanosauriform dinosaur from the Early Cretaceous (Barremian) Yixian Formation of China. The genus contains a single species, ''Ruixinia zhangi''. The ''Ruixinia'' holotype is a partial articulated skeleton with the most complete series of caudal vertebrae known from any Asian titanosauriform. Discovery and naming The ''Ruixinia'' holotype specimen, ELDM EL-J009, was found in sediments of the Yixian Formation in Beipiao, Liaoning Province, northeastern China. This locality is dated to the Barremian age of the Early Cretaceous period. The fossil material consists of a partial articulated skeleton including cervical, dorsal, and sacral vertebrae, caudal vertebrae and associated chevrons, dorsal ribs, and a left ilium, pubis, femur, tibia, astragalus, metatarsal V, and possible pedal phalanx. At the time of its description, the fossil material was still partially embedded in matrix, with only the left side of the bones ...
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Pukyongosaurus
''Pukyongosaurus'' (meaning "Pukyong lizard", after the Pukyong National University) is a genus of titanosauriform dinosaur that lived in South Korea during the Early Cretaceous period ( Aptian - Albian). It may have been closely related to ''Euhelopus'', and is known from a series of vertebrae in the neck and back. The characteristics that were originally used to distinguish this genus have been criticized as being either widespread or too poorly preserved to evaluate, rendering the genus an indeterminate nomen dubium among titanosauriforms. The 2022 study noted that ''Pukyongosaurus'' is probably a somphospondylan. Discovery In 2001, several fragments of a sauropod skeleton were discovered in the Hasandong Formation in Hadong County, South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia, constituting the southern part of the Korean Peninsula and sharing a land border with North Korea. Its western border is formed by the Yellow Sea ...
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