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Thalassodromidae
Thalassodrominae or Thalassodromidae (meaning "sea runners", due to previous misconceptions of skimming behavior; they are now thought to be terrestrial predators) is a group of azhdarchoid pterosaurs from the Cretaceous period. Its traditional members come from Brazil, however, other possible members also come from other places, including the United States, Morocco, and Argentina. Thalassodrominae is considered either to be a subfamily within the pterosaur family Tapejaridae, or as a distinct family, Thalassodromidae, within the clade Neoazhdarchia, closely related to dsungaripterids or azhdarchids. Classification Thalassodrominae traditionally includes only two genera, '' Thalassodromeus'' and ''Tupuxuara'', and was defined to include them and any other descendants of their most recent common ancestor. The classification of thalassodromines is controversial. Some studies, including one by Lü and colleagues in 2008, have found that the thalassodromines are more closely related ...
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Thalassodromeus Sethi
''Thalassodromeus'' is a genus of pterosaur that lived in what is now Brazil during the Early Cretaceous period, about a hundred million years ago. The original skull, discovered in 1983 in the Araripe Basin of northeastern Brazil, was collected in several pieces. In 2002, the skull was made the holotype specimen of ''Thalassodromeus sethi'' by palaeontologists Alexander Kellner and Diogenes de Almeida Campos. The generic name means "sea runner" (in reference to its supposed mode of feeding), and the specific name refers to the Egyptian god Seth due to its crest being supposedly reminiscent of Seth's crown. Other scholars have pointed out that the crest was instead similar to the crown of Amon. A jaw tip was assigned to ''T. sethi'' in 2005, became the basis of the new genus ''Banguela'' in 2014, and assigned back to ''Thalassodromeus'' as the species ''T. oberlii'' in 2018. Another species (''T. sebesensis'') was described in 2015 based on a supposed crest fragment, but this w ...
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Thalassodromeus
''Thalassodromeus'' is a genus of pterosaur that lived in what is now Brazil during the Early Cretaceous period, about a hundred million years ago. The original skull, discovered in 1983 in the Araripe Basin of northeastern Brazil, was collected in several pieces. In 2002, the skull was made the holotype specimen of ''Thalassodromeus sethi'' by palaeontologists Alexander Kellner and Diogenes de Almeida Campos. The generic name means "sea runner" (in reference to its supposed mode of feeding), and the specific name refers to the Egyptian god Seth due to its crest being supposedly reminiscent of Seth's crown. Other scholars have pointed out that the crest was instead similar to the crown of Amon. A jaw tip was assigned to ''T. sethi'' in 2005, became the basis of the new genus '' Banguela'' in 2014, and assigned back to ''Thalassodromeus'' as the species ''T. oberlii'' in 2018. Another species (''T. sebesensis'') was described in 2015 based on a supposed crest fragment, but this ...
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Tupuxuara Leonardii
''Tupuxuara'' is a genus of large, crested, and toothless pterodactyloid pterosaur from the Early Cretaceous period (Albian stage) of what is now the Romualdo Formation of the Santana Group, Brazil, about 125 to 112 million years ago. ''Tupuxuara'' is a close relative of ''Thalassodromeus'', and both form a group that is either called Thalassodrominae (if placed within the family Tapejaridae) or Thalassodromidae (if placed within the clade Neoazhdarchia). Discovery The genus was named and described by Alexander Kellner and Diógenes de Almeida Campos in 1988.Kellner, A.W.A., and Campos, D.A. (1988). "Sobre un novo pterossauro com crista sagital da Bacia do Araripe, Cretaceo Inferior do Nordeste do Brasil. (Pterosauria, Tupuxuara, Cretaceo, Brasil)." ''Anais da Academia Brasileira de Ciências'', 60: 459–469. n Portuguese/ref> The type species is ''Tupuxuara longicristatus''. The generic name refers to a familiar spirit from the mythology of the Tupi. The specific n ...
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Tupuxuara
''Tupuxuara'' is a genus of large, crested, and toothless pterodactyloid pterosaur from the Early Cretaceous period (Albian stage) of what is now the Romualdo Formation of the Santana Group, Brazil, about 125 to 112 million years ago. ''Tupuxuara'' is a close relative of ''Thalassodromeus'', and both form a group that is either called Thalassodrominae (if placed within the family Tapejaridae) or Thalassodromidae (if placed within the clade Neoazhdarchia). Discovery The genus was named and described by Alexander Kellner and Diógenes de Almeida Campos in 1988.Kellner, A.W.A., and Campos, D.A. (1988). "Sobre un novo pterossauro com crista sagital da Bacia do Araripe, Cretaceo Inferior do Nordeste do Brasil. (Pterosauria, Tupuxuara, Cretaceo, Brasil)." ''Anais da Academia Brasileira de Ciências'', 60: 459–469. n Portuguese/ref> The type species is ''Tupuxuara longicristatus''. The generic name refers to a familiar spirit from the mythology of the Tupi. The specific n ...
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Banguela
''Banguela'' is a genus of azhdarchoid pterosaur from the Early Cretaceous period (Albian stage) of what is now Brazil. Only one species is known, ''Banguela oberlii''. Discovery and naming The Swiss collector Urs Oberli acquired a pterosaur jaw fragment from the Chapada do Araripe. In 2005, this was described by André Jacques Veldmeijer and colleagues, and referred to '' Thalassodromeus sethi''.Veldmeijer, A.J., Signore, M. & Meijer, H.J.M., 2005, "Description of two pterosaur (Pterodactyloidea) mandibles from the lower Cretaceous Santana Formation, Brazil", ''DEINSEA'' 11: 67–86 In 2014 it was named and described by Jaime Headden and Hebert Bruno Nascimento Campos as a separate genus ''Banguela'', with the type species ''Banguela oberlii''. The generic name is a Brazilian Portuguese word for "toothless one", especially used as an affectionate term for elderly women. The specific name honors Oberli. The holotype, NMSG SAO 251093, was probably found in the Romualdo Form ...
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Pterosaurs
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or ' rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insecti ...
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Neoazhdarchia
Azhdarchoidea (or azhdarchoids) is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea. Pterosaurs belonging to this group lived throughout the Early and Late Cretaceous periods, with one tentative member, '' Tendaguripterus'', that lived in the Late Jurassic period. The largest azhdarchoids include members of the family Azhdarchidae, examples of these are ''Quetzalcoatlus'', ''Hatzegopteryx'', and '' Arambourgiania''. The Azhdarchoidea has been recovered as either closely related to the Ctenochasmatoidea, as the sister taxon of the Pteranodontoidea within the Ornithocheiroidea, or within the Tapejaroidea, which in turn was also within the Ornithocheiroidea. Classification Azhdarchoidea was given a phylogenetic definition by David Unwin in 2003. Unwin defined the group as the most recent common ancestor of ''Quetzalcoatlus'' and '' Tapejara'', and all its descendants.Unwin, D. M., (2003). "On the phylogeny and evolution ...
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Azhdarchoid
Azhdarchoidea (or azhdarchoids) is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea. Pterosaurs belonging to this group lived throughout the Early and Late Cretaceous periods, with one tentative member, ''Tendaguripterus'', that lived in the Late Jurassic period. The largest azhdarchoids include members of the family Azhdarchidae, examples of these are ''Quetzalcoatlus'', ''Hatzegopteryx'', and ''Arambourgiania''. The Azhdarchoidea has been recovered as either closely related to the Ctenochasmatoidea, as the sister taxon of the Pteranodontoidea within the Ornithocheiroidea, or within the Tapejaroidea, which in turn was also within the Ornithocheiroidea. Classification Azhdarchoidea was given a phylogenetic definition by David Unwin in 2003. Unwin defined the group as the most recent common ancestor of ''Quetzalcoatlus'' and '' Tapejara'', and all its descendants.Unwin, D. M., (2003). "On the phylogeny and evolutionar ...
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Lacusovagus
''Lacusovagus'' (meaning "lake wanderer") is a genus of azhdarchoid pterodactyloid pterosaur from the Lower Cretaceous of Brazil. It is based on SMNK PAL 4325, a partial upper jaw comprising sections of the skull in front of the eyes. This specimen was found in rocks of the Early Cretaceous-age (probably Aptian stage, about 120 million years ago) Nova Olinda Member of the Crato Formation. The skull was long, and unusually wide. The section in front of the combined nasal-antorbital fenestra was relatively short. Also unusual was the combination of its toothless jaws and no bony head crest. ''Lacusovagus'' was described in 2008 by Mark Witton. The type species is ''L. magnificens'', meaning "grand lake wanderer", in reference to its large size—it is currently the largest pterosaur known from the Crato Formation with an estimated wingspan of approximately and a body mass of . ''Lacusovagus'' shares many characteristics with the basal azhdarchoid family Chaoyangopteridae, ...
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Azhdarchoidea
Azhdarchoidea (or azhdarchoids) is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea. Pterosaurs belonging to this group lived throughout the Early and Late Cretaceous periods, with one tentative member, '' Tendaguripterus'', that lived in the Late Jurassic period. The largest azhdarchoids include members of the family Azhdarchidae, examples of these are ''Quetzalcoatlus'', ''Hatzegopteryx'', and ''Arambourgiania''. The Azhdarchoidea has been recovered as either closely related to the Ctenochasmatoidea, as the sister taxon of the Pteranodontoidea within the Ornithocheiroidea, or within the Tapejaroidea, which in turn was also within the Ornithocheiroidea. Classification Azhdarchoidea was given a phylogenetic definition by David Unwin in 2003. Unwin defined the group as the most recent common ancestor of ''Quetzalcoatlus'' and '' Tapejara'', and all its descendants.Unwin, D. M., (2003). "On the phylogeny and evolutio ...
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Alanqa
''Alanqa'' is a genus of pterodactyloid pterosaur from the Late Cretaceous period (Cenomanian stage) of what is now the Kem Kem Beds of southeastern Morocco. The name ''Alanqa'' comes from the Arabic language, Arabic word العنقاء ''al-Anqa, ‘Anqā’'', for a mythical bird of Arabian culture. Discovery Aided by local villagers, a team of paleontologists had been excavating at several locations in the Kem Kem Beds during April, and November to December 2008, uncovering remains of several different pterosaurs. The material was fragmentary, and the type locality for ''Alanqa'' is Aferdou N'Chaft, near the village of Begaa and to the north-east of Taouz. Description ''Alanqa'' is known only from five fragments of the front upper and lower jaws, and possibly a neck vertebra, representing the single type species ''Alanqa saharica''. Two of these fragments were first described, but not named, by Peter Wellnhofer, Wellnhofer and Éric Buffetaut, Buffetaut in 1999, and referre ...
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Aerotitan
''Aerotitan'' is a genus of large azhdarchid pterosaur known from the Late Cretaceous period (Maastrichtian stage) of what is now the Allen Formation of the Neuquén Basin in northern Patagonia, Argentina. Discovery and naming The type species ''Aerotitan sudamericanus'' was named and described in 2012 by Fernando Novas, Martin Kundrat, Federico Agnolín, Martin Ezcurra, Per Erik Ahlberg, Marcelo Isasi, Alberto Arriagada and Pablo Chafrat. The generic name is derived from Greek ἀήρ, ''aer'', "air", and Titan, in reference to the fact the species represents a large flying reptile. The specific name refers to its provenance from South America. The holotype, MPCN-PV 0054, has been recovered near the Bajo de Arriagada site, in Patagonia, from a layer of the upper Allen Formation. It consists, according to the original description, of a partial rostrum with a preserved length of . This snout is elongated and transversely compressed and the jaws are toothless. The wingsp ...
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