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Tiphioidea
Tiphioidea is a suggested superfamily of stinging wasps in the order Hymenoptera. There are three families in Tiphioidea, Bradynobaenidae, Tiphiidae, and Sierolomorphidae. Recent research in molecular phylogenetics has resulted in the reorganization of the infraorder Aculeata, which now contains eight superfamilies: Apoidea, Chrysidoidea, Formicoidea, Pompiloidea, Scolioidea, Tiphioidea, Thynnoidea, and Vespoidea Vespoidea is a superfamily of wasps in the order Hymenoptera, although older taxonomic schemes may vary in this categorization, particularly in whether to recognize the superfamilies Scolioidea (for scoliid wasps) or Formicoidea (for ants) .... References Further reading * * * External links * Hymenoptera Apocrita superfamilies {{hymenoptera-stub ...
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Vespoidea
Vespoidea is a superfamily of wasps in the order Hymenoptera, although older taxonomic schemes may vary in this categorization, particularly in whether to recognize the superfamilies Scolioidea (for scoliid wasps) or Formicoidea (for ants). Vespoidea includes wasps with a large variety of lifestyles: eusocial, social, and solitary habits, predators, scavengers, parasitoids, and some herbivores. Description Vespoid wasp females have antennae with 10 flagellomeres, while males have 11 flagellomeres. The edge of the pronotum reaches or passes the tegula. Many species display some level of sexual dimorphism. Most species have fully developed wings, but some have reduced or absent wings in one or both sexes. As in other Aculeata, only the females are ever capable of stinging. Phylogenetics and taxonomy Research based on four nuclear genes (elongation factor-1α F2 copy, long-wavelength rhodopsin, wingless and the D2–D3 regions of 28S ribosomal RNA—2700 bp in total) su ...
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Bradynobaenidae
The Bradynobaenidae are a family of wasps similar to the Mutillidae, differing most visibly in the presence, in females, of a suture separating the pronotum from the mesonotum. These species are often found in arid regions. Recent classifications (beginning in 2008) remove two of the five constituent genera, both from the New World, to a separate family Chyphotidae, thus restricting true bradynobaenids to the Old World The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by th .... Genera * '' Apterogyna'' Latreille, 1809 * '' Gynecaptera'' Skorikov, 1935 * '' Bradynobaenus'' Spinola, 1851 * '' Macroocula'' Panfilov, 1954 * '' Micatagla'' Argaman, 1994 (at least 54 living species per Gadallah, et al., 2019) Genera placed in Chyphotidae * '' Chyphotes'' Blake, 1886 * '' Typhoctes'' Ashme ...
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Sierolomorphidae
The Sierolomorphidae are a family of 13 extant species of wasps, in the genera '' Sierolomorpha'' and '' Proscleroderma'', mostly found in the Northern Hemisphere. They are rare and very little is known of their biology. A fossil species '' Loreisomorpha nascimbenei'' has also been placed in the family. The coxa (basal segment of the leg) of the hind and midlegs are next to each other, and the hindwing does not have claval or jugal lobes. The first metasomal segment does not have a true node, but can appear like that of the ants. The metasomal sternum of the first segment is separated from the second by a constriction. Sexual dimorphism varies among species from slight to marked, with both males and females having wings, but females are sometimes wingless. Adults are predominantly dark brown or black in colour. They are solitary and the larvae are suspected to be ectoparasitoid In evolutionary ecology, a parasitoid is an organism that lives in close association with its hos ...
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Aculeata
Aculeata is a subclade of Hymenoptera containing ants, bees, and stinging wasps. The name is a reference to the defining feature of the group, which is the modification of the ovipositor into a stinger. However, many members of the group cannot sting, either retaining the ovipositor, or having lost it altogether. A large part of the clade is parasitic. This group includes all of the eusocial Hymenopterans. It is theorized that the possession of a venomous sting was important in the repeated evolution of eusociality within Hymenoptera. The oldest aculeates are known from the Late Jurassic Karabastau Formation of Kazakhstan, represented by the family Bethylonymidae, which may be para or polyphyletic. Classification The use of the name Aculeata has a long history at the rank of infraorder or division. The Aculeata are a monophyletic, or good natural group, containing all the descendants of a single common ancestor. The Aculeata are therefore maintained as a taxon In bio ...
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Hymenoptera
Hymenoptera is a large order of insects, comprising the sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are parasitic. Females typically have a special ovipositor for inserting eggs into hosts or places that are otherwise inaccessible. This ovipositor is often modified into a stinger. The young develop through holometabolism (complete metamorphosis)—that is, they have a wormlike larval stage and an inactive pupal stage before they mature. Etymology The name Hymenoptera refers to the wings of the insects, but the original derivation is ambiguous. All references agree that the derivation involves the Ancient Greek πτερόν (''pteron'') for wing. The Ancient Greek ὑμήν (''hymen'') for membrane provides a plausible etymology for the term because species in this order have membranous wings. However, a key characteristic of this order is that the hindwings are co ...
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Tiphiidae
The Tiphiidae (also known as the tiphiid wasps) are a family of large, solitary wasps whose larvae are parasitoids of various beetle larvae, especially those in the superfamily Scarabaeoidea. Until recently, this family contained several additional subfamilies, but multiple studies have independently confirmed that these comprise a separate lineage, and are now classified in the family Thynnidae. The females of some Brachycistidinae are wingless, and hunt ground-dwelling (fossorial) beetle larvae. The prey is paralysed with the female's sting, and an egg is lain on it so the wasp larva has a ready supply of food. As some of the ground-dwelling scarab species attacked by tiphiids are pests, some of these wasps are considered beneficial as biological control agents. Taxonomy Tiphiid genera are classified as follows: Subfamily Brachycistidinae *'' Acanthetropis'' Wasbauer, 1958 *'' Brachycistellus'' Baker, 1907 *'' Brachycistina'' Malloch, 1926 *'' Brachycistis'' Fox, 1 ...
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Tiphia P1280629a
''Tiphia'' is a genus of wasps belonging to the family Tiphiidae The Tiphiidae (also known as the tiphiid wasps) are a family of large, solitary wasps whose larvae are parasitoids of various beetle larvae, especially those in the superfamily Scarabaeoidea. Until recently, this family contained several additiona ... subfamily Tiphiinae. They feed on soil-inhabiting scarab beetle larvae. The species ''Tiphia vernalis'' (Spring Tiphia) has been introduced in the United States from Korea and China in 1925 to combat the Japanese beetle (''Popillia japonica''). Species * ''Tiphia abnormis'' Eversmann, 1849 * ''Tiphia alishana'' Ishikawa, 1967 * ''Tiphia ami'' Tsuneki, 1986 * ''Tiphia annulata'' Fabricius, 1793 * ''Tiphia antigae'' Tournier, 1901 * ''Tiphia arthroxantha'' Boni Bartalucci, 2011 * ''Tiphia austriaca'' Tournier, 1889 * ''Tiphia bexar'' Nagy, 1967 * ''Tiphia brevala'' Zhang, 1989 * ''Tiphia bunun'' Tsuneki, 1986 * Tiphia chareshi * ''Tiphia changi'' Tsuneki, 1986 * ' ...
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Apoidea
The superfamily Apoidea is a major group within the Hymenoptera, which includes two traditionally recognized lineages, the "sphecoid" wasps, and the bees. Molecular phylogeny demonstrates that the bees arose from within the traditional "Crabronidae", so that grouping is paraphyletic, and this has led to a reclassification to produce monophyletic families.Manuela Sann, Oliver Niehuis, Ralph S. Peters, Christoph Mayer, Alexey Kozlov, Lars Podsiadlowski, Sarah Bank, Karen Meusemann, Bernhard Misof, Christoph Bleidorn and Michael Ohl (2018) Phylogenomic analysis of Apoidea sheds new light on the sister group of bees. ''BMC Evolutionary Biology'' 18:71. doi:10.1186/s12862-018-1155-8 Nomenclature Bees appear in recent classifications to be a specialized lineage of " crabronid" wasps that switched to the use of pollen and nectar as larval food, rather than insect prey; this makes the traditional "Crabronidae" a paraphyletic group. Accordingly, bees and sphecoids are now all grouped toge ...
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Chrysidoidea
The superfamily Chrysidoidea is a very large cosmopolitan group (some 6,000 described species, and many more undescribed) , all of which are parasitoids or cleptoparasites of other insects. There are three large, common families (Bethylidae, Chrysididae, and Dryinidae) and four small, rare families ( Embolemidae, Plumariidae, Sclerogibbidae, and Scolebythidae). Most species are small (7 mm or less), almost never exceeding 15 mm. This superfamily is traditionally considered to be the basal taxon within the Aculeata, and, as such, some species can sting, though the venom is harmless to humans. Members of the families Dryinidae and Embolemidae are the only parasitoids among the Hymenoptera to have a life cycle in which the wasp larva begins its life inside the body of the host, and then later forms a sac (called a ''thylacium'') that protrudes out of the host's abdomen. The closely related family Sclerogibbidae contains more traditional ectoparasitoids, attacking t ...
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Formicoidea
Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22,000 species have been classified. They are easily identified by their geniculate (elbowed) antennae and the distinctive node-like structure that forms their slender waists. Ants form colonies that range in size from a few dozen predatory individuals living in small natural cavities to highly organised colonies that may occupy large territories and consist of millions of individuals. Larger colonies consist of various castes of sterile, wingless females, most of which are workers (ergates), as well as soldiers (dinergates) and other specialised groups. Nearly all ant colonies also have some fertile males called "drones" and one or more fertile females called "queens" ( gynes). The colonies are described as superorganisms because the ants ap ...
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Pompiloidea
Pompiloidea is a superfamily that includes spider wasps and velvet ants, among others. in the order Hymenoptera. There are 4 families in Pompiloidea. Families These four families belong to the superfamily Pompiloidea: * Mutillidae (velvet ants) * Myrmosidae (myrmosid wasps) * Pompilidae (spider wasps) * Sapygidae The Sapygidae are a family of solitary kleptoparasitic aculeate wasps. They are generally black wasps, similar in appearance to some Tiphiidae or Thynnidae, with white or yellow markings developed to various degrees. The female oviposits her ... (sapygid wasps) The extinct family Burmusculidae, known from Cretaceous amber, is also placed here.Longfeng Li; Alexandr P. Rasnitsyn; Chungkun Shih; Daqing Li; Dong Ren (2020). "Two new rare wasps (Hymenoptera: Apocrita: Panguidae and Burmusculidae) from mid-Cretaceous amber of northern Myanmar". Cretaceous Research. 109: Article 104220. doi:10.1016/j.cretres.2019.104220 References Further reading * * * * E ...
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Scolioidea
The Scoliidae, the scoliid wasps, are a family of about 560 species found worldwide. They tend to be black, often marked with yellow or orange, and their wing tips are distinctively corrugated. Males are more slender and elongated than females, with significantly longer antennae, but the sexual dimorphism is not as apparent as in the Tiphiidae. Biology Scoliid wasps are solitary parasitoids of scarab beetle larvae. Female scoliids burrow into the ground in search of these larvae and then use their sting to paralyze them. They will sometimes excavate a chamber and move the paralyzed beetle larva into it before depositing an egg. Scoliid wasps act as important biocontrol agents, as many of the beetles they parasitize are pests, including the Japanese beetle. Male scoliids patrol territories, ready to mate with females emerging from the ground. Adult wasps may be minor pollinators of some plants and can be found on many wildflowers in the late summer. Scoliidae also has at least ...
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