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Stigmaria
''Stigmaria'' is a form taxon for common fossils found in Carboniferous rocks. They represent the underground rooting structures of coal forest lycopsid trees such as '' Sigillaria'' and '' Lepidodendron''. These swamp forest trees grew to 50 meters and were anchored by an extensive network of branching underground structures with "rootlets" attached to them. Analysis of the morphology and anatomy of these stigmarian systems suggests they were shoot-like and so they are called rhizomes or rhizophores. The stigmarian rhizomes are typically covered with a spiral pattern of circular scars where "rootlets" were attached. Since the stigmarian systems are shoot-like, these "rootlets" may be modified leaves, adapted to serve the function of roots. However, some paleontologists argue that the "rootlets" were true roots, with a complex branching structure and root hairs, comparable to the roots of the closest living relative of ''Lepidodendron'', the quillworts (genus ''Isoetes ''I ...
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Stigmaria Mcr1
''Stigmaria'' is a form taxon (botany), form taxon for common fossils found in Carboniferous rocks. They represent the underground rooting structures of coal forest lycopsid trees such as ''Sigillaria'' and ''Lepidodendron''. These swamp forest trees grew to 50 meters and were anchored by an extensive network of branching underground structures with "rootlets" attached to them. Analysis of the morphology and anatomy of these stigmarian systems suggests they were shoot-like and so they are called rhizomes or rhizophores. The stigmarian rhizomes are typically covered with a spiral pattern of circular scars where "rootlets" were attached. Since the stigmarian systems are shoot-like, these "rootlets" may be modified leaves, adapted to serve the function of roots. However, some paleontologists argue that the "rootlets" were true roots, with a complex branching structure and root hairs, comparable to the roots of the closest living relative of ''Lepidodendron'', the quillworts (genus ...
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Carboniferous
The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carboniferous'' means "coal-bearing", from the Latin '' carbō'' ("coal") and '' ferō'' ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern 'system' names, it was coined by geologists William Conybeare and William Phillips in 1822, based on a study of the British rock succession. The Carboniferous is often treated in North America as two geological periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial animal life was well established by the Carboniferous Period. Tetrapods (four limbed vertebrates), which had originated from lobe-finned fish during the preceding Devonian, became pentadactylous in and diversified during the Carboniferous, including early amphibian lin ...
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Carboniferous Plants
The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carboniferous'' means "coal-bearing", from the Latin '' carbō'' ("coal") and '' ferō'' ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern 'system' names, it was coined by geologists William Conybeare and William Phillips in 1822, based on a study of the British rock succession. The Carboniferous is often treated in North America as two geological periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial animal life was well established by the Carboniferous Period. Tetrapods (four limbed vertebrates), which had originated from lobe-finned fish during the preceding Devonian, became pentadactylous in and diversified during the Carboniferous, including early amphibian li ...
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Lepidodendron
''Lepidodendron'' is an extinct genus of primitive vascular plants belonging to the family Lepidodendraceae, part of a group of Lycopodiopsida known as scale trees or arborescent lycophytes, related to quillworts and lycopsids (club mosses). They were part of the coal forest flora. They sometimes reached heights of , and the trunks were often over in diameter. They thrived during the Carboniferous Period (358.9 to 298.9 million years ago). Sometimes erroneously called "giant club mosses", the genus was actually more closely related to modern quillworts than to modern club mosses. Within the form classification system used within paleobotany, ''Lepidodendron'' is both used for the whole plant as well as specifically the stems and leaves. Etymology The name ''Lepidodendron'' comes from the Greek λεπίς ', scale, and δένδρον ''dendron'', tree. Growth During the early stages of growth, ''Lepidodendron'' grew as single, unbranched trunk, with leaves growing out of ...
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Sigillaria
''Sigillaria'' is a genus of extinct, spore-bearing, arborescent (tree-like) plants. It was a lycopodiophyte, and is related to the lycopsids, or club-mosses, but even more closely to quillworts, as was its associate ''Lepidodendron''. Fossil records This genus is known in the fossil records from as early as the Middle Devonian or the Late Carboniferous period but dwindled to extinction in the Early Permian period (age range: from 383.7 to 254.0 million years ago). Fossils are found in Great Britain, United States, Canada, China, Korea, Tanzania and Zimbabwe. Species Species within this genus include: *''S.alveolaris'' Brongniart (1828) *''S.barbata'' Weiss (1887) *''S.bicostata'' Weiss (1887) *''S.boblayi'' Brongniart (1828) *''S.brardii'' Brongniart (1828) *''S.cancriformis'' Weiss (1887) *''S.cristata'' Sauveur (1848) *''S.cumulata'' Weiss (1887) *''S.davreuxii'' Brongniart (1828) *''S.densifolia'' Brongniart (1836) *''S.elegans'' Sternberg (1825) *''S.elongata'' Brongn ...
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Lycopsid Joggins Mcr1
Lycopodiopsida is a class of vascular plants known as lycopods, lycophytes or other terms including the component lyco-. Members of the class are also called clubmosses, firmosses, spikemosses and quillworts. They have dichotomously branching stems bearing simple leaves called microphylls and reproduce by means of spores borne in sporangia on the sides of the stems at the bases of the leaves. Although living species are small, during the Carboniferous, extinct tree-like forms formed huge forests that dominated the landscape and contributed to coal deposits. The nomenclature and classification of plants with microphylls varies substantially among authors. A consensus classification for extant (living) species was produced in 2016 by the Pteridophyte Phylogeny Group (PPG I), which places them all in the class Lycopodiopsida, which includes the classes Isoetopsida and Selaginellopsida used in other systems. (See Table 2.) Alternative classification systems have used ranks f ...
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Adolphe-Théodore Brongniart
Adolphe-Théodore Brongniart () FRS FRSE FGS (14 January 1801 – 18 February 1876) was a French botanist. He was the son of the geologist Alexandre Brongniart and grandson of the architect, Alexandre-Théodore Brongniart. Brongniart's pioneering work on the relationships between extinct and existing plants has earned him the title of father of paleobotany. His major work on plant fossils was his ' (1828–37). He wrote his dissertation on the Buckthorn family (Rhamnaceae), an extant family of flowering plants, and worked at the Muséum national d'Histoire naturelle in Paris until his death. In 1851, he was elected a foreign member of the Royal Swedish Academy of Sciences. This botanist is denoted by the author abbreviation when citing a botanical name. Brongniart's works Brongniart was an indefatigable investigator and a prolific writer of books and memoirs. As early as 1822 he published a paper on the classification and distribution of fossil plants. This was followed ...
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Form Taxon (botany)
Form classification is the classification of organisms based on their morphology, which does not necessarily reflect their biological relationships. Form classification, generally restricted to palaeontology, reflects uncertainty; the goal of science is to move "form taxa" to biological taxa whose affinity is known. Form taxonomy is restricted to fossils that preserve too few characters for a conclusive taxonomic definition or assessment of their biological affinity, but whose study is made easier if a binomial name is available by which to identify them. The term "form classification" is preferred to "form taxonomy"; taxonomy suggests that the classification implies a biological affinity, whereas form classification is about giving a name to a group of morphologically-similar organisms that may not be related. A "parataxon" (not to be confused with parataxonomy), or "sciotaxon" (Gr. "shadow taxon"), is a classification based on incomplete data: for instance, the larval stage ...
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Coal Forest
Coal forests were the vast swathes of wetlands that covered much of the Earth's tropical land areas during the late Carboniferous ( Pennsylvanian) and Permian times.Cleal, C. J. & Thomas, B. A. (2005). "Palaeozoic tropical rainforests and their effect on global climates: is the past the key to the present?" ''Geobiology'', ''3'', p. 13-31. As vegetable matter from these forests decayed, enormous deposits of peat accumulated, which later changed into coal. Much of the carbon in the peat deposits produced by coal forests came from photosynthetic splitting of existing carbon dioxide, which released the accompanying split-off oxygen into the atmosphere. This process may have greatly increased the oxygen level, possibly as high as about 35%, making the air more easily breathable by animals with inefficient respiratory systems, as indicated by the size of ''Meganeura'' compared to modern dragonflies. Coal forests covered tropical Euramerica (Europe, eastern North America, northwestern ...
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Lycopsid
Lycopodiopsida is a class of vascular plants known as lycopods, lycophytes or other terms including the component lyco-. Members of the class are also called clubmosses, firmosses, spikemosses and quillworts. They have dichotomously branching stems bearing simple leaves called microphylls and reproduce by means of spores borne in sporangia on the sides of the stems at the bases of the leaves. Although living species are small, during the Carboniferous, extinct tree-like forms formed huge forests that dominated the landscape and contributed to coal deposits. The nomenclature and classification of plants with microphylls varies substantially among authors. A consensus classification for extant (living) species was produced in 2016 by the Pteridophyte Phylogeny Group (PPG I), which places them all in the class Lycopodiopsida, which includes the classes Isoetopsida and Selaginellopsida used in other systems. (See Table 2.) Alternative classification systems have used ranks fr ...
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Permian
The Permian ( ) is a geologic period and System (stratigraphy), stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the Perm Governorate, region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the Sauropsida, sauropsids (reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditi ...
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Rhizome
In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow horizontally. The rhizome also retains the ability to allow new shoots to grow upwards. A rhizome is the main stem of the plant that runs underground horizontally. A stolon is similar to a rhizome, but a stolon sprouts from an existing stem, has long internodes, and generates new shoots at the end, such as in the strawberry plant. In general, rhizomes have short internodes, send out roots from the bottom of the nodes, and generate new upward-growing shoots from the top of the nodes. A stem tuber is a thickened part of a rhizome or stolon that has been enlarged for use as a storage organ. In general, a tuber is high in starch, e.g. the potato, which is a modified stolon. The term "tuber" is often used imprecisely and is sometimes applied ...
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