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Sauropsid
Sauropsida ( Greek for "lizard faces") is a clade of amniotes, broadly equivalent to the class Reptilia, though typically used in a broader sense to also include extinct stem-group relatives of modern reptiles and birds (which, as theropod dinosaurs, are nested within reptiles as more closely related to crocodilians than to lizards or turtles).Gauthier J.A. (1994): ''The diversification of the amniotes''. In: D.R. Prothero and R.M. Schoch (ed.) Major Features of Vertebrate Evolution: 129–159. Knoxville, Tennessee: The Paleontological Society. The most popular definition states that Sauropsida is the sibling taxon to Synapsida, the other clade of amniotes which includes mammals as its only modern representatives. Although early synapsids have historically been referred to as "mammal-like reptiles", all synapsids are more closely related to mammals than to any modern reptile. Sauropsids, on the other hand, include all amniotes more closely related to modern reptiles than to mam ...
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Parareptilia
Parareptilia ("near-reptiles") is an extinct group of basal sauropsids (" reptiles"), traditionally considered the sister taxon to Eureptilia (the group that likely contains all living reptiles and birds). Parareptiles first arose near the end of the Carboniferous period and achieved their highest diversity during the Permian period. Several ecological innovations were first accomplished by parareptiles among reptiles. These include the first reptiles to return to marine ecosystems (mesosaurs), the first bipedal reptiles ( bolosaurids such as '' Eudibamus''), the first reptiles with advanced hearing systems ( nycteroleterids and others), and the first large herbivorous reptiles (the pareiasaurs). The only parareptiles to survive into the Triassic period were the procolophonoids, a group of small generalists, omnivores, and herbivores. The largest family of procolophonoids, the procolophonids, rediversified in the Triassic, but subsequently declined and became extinct by t ...
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Mesosaur
Mesosaurs ("middle lizards") were a group of small aquatic reptiles that lived during the early Permian period ( Cisuralian), roughly 299 to 270 million years ago. Mesosaurs were the first known aquatic reptiles, having apparently returned to an aquatic lifestyle from more terrestrial ancestors. It is uncertain which and how many terrestrial traits these ancestors displayed; recent research cannot establish with confidence if the first amniotes were fully terrestrial, or only amphibious. Most authors consider mesosaurs to have been aquatic, although adult animals may have been amphibious, rather than completely aquatic, as indicated by their moderate skeletal adaptations to a semiaquatic lifestyle.Pablo Nuñez Demarco et al. Was Mesosaurus a Fully Aquatic Reptile? Front. Ecol. Evol, published online July 27, 2018; doi: 10.3389/fevo.2018.00109 Similarly, their affinities are uncertain; they may have been among the most basal sauropsids or among the most basal parareptiles (in ...
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Mesosauria
Mesosaurs ("middle lizards") were a group of small aquatic reptiles that lived during the early Permian period (Cisuralian), roughly Geologic time scale, 299 to 270 million years ago. Mesosaurs were the first known aquatic reptiles, having apparently returned to an aquatic lifestyle from more terrestrial ancestors. It is uncertain which and how many terrestrial traits these ancestors displayed; recent research cannot establish with confidence if the first amniotes were fully terrestrial, or only amphibious. Most authors consider mesosaurs to have been aquatic, although adult animals may have been amphibious, rather than completely aquatic, as indicated by their moderate skeletal adaptations to a semiaquatic lifestyle.Pablo Nuñez Demarco et al. Was Mesosaurus a Fully Aquatic Reptile? Front. Ecol. Evol, published online July 27, 2018; doi: 10.3389/fevo.2018.00109 Similarly, their affinities are uncertain; they may have been among the most basal sauropsids or among the most basal p ...
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Parareptile
Parareptilia ("near-reptiles") is an extinct group of Basal (phylogenetics), basal Sauropsida, sauropsids ("Reptile, reptiles"), traditionally considered the sister taxon to Eureptilia (the group that likely contains all living reptiles and birds). Parareptiles first arose near the end of the Carboniferous, Carboniferous period and achieved their highest diversity during the Permian, Permian period. Several ecological innovations were first accomplished by parareptiles among reptiles. These include the first reptiles to return to marine ecosystems (mesosaurs), the first Bipedalism, bipedal reptiles (Bolosauridae, bolosaurids such as ''Eudibamus''), the first reptiles with advanced hearing systems (Nycteroleteridae, nycteroleterids and others), and the first large herbivorous reptiles (the pareiasaurs). The only parareptiles to survive into the Triassic, Triassic period were the Procolophonoidea, procolophonoids, a group of small generalists, omnivores, and herbivores. The largest f ...
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Carboniferous
The Carboniferous ( ) is a Geologic time scale, geologic period and System (stratigraphy), system of the Paleozoic era (geology), era that spans 60 million years, from the end of the Devonian Period Ma (million years ago) to the beginning of the Permian Period, Ma. It is the fifth and penultimate period of the Paleozoic era and the fifth period of the Phanerozoic eon (geology), eon. In North America, the Carboniferous is often treated as two separate geological periods, the earlier Mississippian (geology), Mississippian and the later Pennsylvanian (geology), Pennsylvanian. The name ''Carboniferous'' means "coal-bearing", from the Latin ("coal") and ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern "system" names, it was coined by geologists William Conybeare (geologist), William Conybeare and William Phillips (geologist), William Phillips in 1822, based on a study of the British rock succession. Carboniferous is the per ...
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Recumbirostra
Recumbirostra is a clade of tetrapods which lived during the Carboniferous and Permian periods. They are thought to have had a fossorial (burrowing) lifestyle and the group includes both short-bodied and long-bodied snake-like forms. At least one species, the long-bodied molgophid '' Nagini mazonense,'' lost its forelimbs entirely. Recumbirostra includes the families Pantylidae, Gymnarthridae, Ostodolepidae, Rhynchonkidae and Brachystelechidae, with additional families such as Microbrachidae and Molgophidae being included by some authors. Brachystelechidae and Molgophidae have also been grouped together in the suggested clade Chthonosauria. Recumbirostra was erected as a clade in 2007 to include many of the taxa traditionally grouped in " Microsauria", which has since been shown to be a paraphyletic or polyphyletic grouping. Like other "microsaurs", the recumbirostrans have traditionally been considered to be members of the subclass Lepospondyli; however, many phylogen ...
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Pennsylvanian (geology)
The Pennsylvanian ( , also known as Upper Carboniferous or Late Carboniferous) is, on the International Commission on Stratigraphy, ICS geologic timescale, the younger of two period (geology), subperiods of the Carboniferous Period (or the upper of two system (stratigraphy), subsystems of the Carboniferous System). It lasted from roughly . As with most other geochronology, geochronologic units, the stratum, rock beds that define the Pennsylvanian are well identified, but the exact date of the start and end are uncertain by a few hundred thousand years. The Pennsylvanian is named after the U.S. state of Pennsylvania, where the coal Bed (geology), beds of this age are widespread. The division between Pennsylvanian and Mississippian (geology), Mississippian comes from North American stratigraphy. In North America, where the early Carboniferous beds are primarily marine limestones, the Pennsylvanian was in the past treated as a full-fledged geologic period between the Mississippian ...
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Captorhinidae
Captorhinidae is an extinct family of tetrapods, traditionally considered primitive Reptile, reptiles, known from the late Carboniferous to the Late Permian. They had a cosmopolitan distribution across Pangea. Description Captorhinids are a clade of small to very large lizard-like animals that date from the Late Carboniferous through the Permian. Their skulls were much stronger than those of their relatives, the protorothyridids, and had teeth that were better able to deal with tough plant material. The postcranial skeleton is similar to those of seymouriamorphs and diadectomorphs; these animals were grouped together with the captorhinids in the order Cotylosauria as the first reptiles in the early 20th century, but are now usually regarded as Stem group, stem-amniotes no closer to reptiles than to mammals. Captorhinids have broad, robust skulls that are generally triangular in shape when seen in dorsal view. The premaxillae are characteristically downturned. The largest captorh ...
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Procolophonia
Procolophonia is an extinct suborder (clade) of herbivorous reptiles that lived from the Middle Permian till the end of the Triassic period. They were originally included as a suborder of the Cotylosauria (later renamed Captorhinida Carroll 1988) but are now considered a clade of Parareptilia. They are closely related to other generally lizard-like Permian reptiles such as the Millerettidae, Bolosauridae, Acleistorhinidae, and Lanthanosuchidae, all of which are included under the Anapsida or "Parareptiles" (as opposed to the Eureptilia). Classification There are two main groups of Procolophonia, the small, lizard-like Procolophonoidea, and the Pareiasauroidea, which include the large, armoured Pareiasauridae. According to the traditional classification of Carroll 1988 as well as phylogenetic analyses of 2012, smaller groups like Rhipaeosauridae (now a synonym of Nycteroleteridae) and Sclerosauridae are classified with the pareiasaurs and with the procolophonids, ...
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Australothyris
''Australothyris'' is an extinct genus of basal procolophonomorph Parareptilia, parareptile known from the Middle Permian (middle Capitanian stage) of Tapinocephalus Assemblage Zone, ''Tapinocephalus'' Assemblage Zone, South Africa. The type and only known species is ''Australothyris smithi''. As the most basal member of Procolophonomorpha, ''Australothyris'' helped to contextualize the origin of this major parareptile subgroup. It has been used to support the hypotheses that procolophonomorphs originated in Gondwana and ancestrally possess Infratemporal fenestra, temporal fenestrae, due to its large and fully enclosed temporal fenestra and South African heritage. It also possessed several unique features, including a high tooth number, long Postfrontal bone, postfrontal, small interpterygoid vacuity, and a specialized interaction between the stapes and Quadrate bone, quadrate. Discovery Australothyris is known from a single specimen discovered at the Beukesplaas farm by Robert ...
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Bolosauria
Procolophonomorpha is an Order (biology), order containing most parareptiles. Many papers have applied various definitions to the name, though most of these definitions have since been considered synonymous with modern parareptile Clade, clades such as Ankyramorpha and Procolophonia. The current definition of Procolophonomorpha, as defined by Modesto, Scott, & Reisz (2009), is that of as a stem-based group containing ''Procolophon'' and all taxa more closely related to it than to ''Milleretta''. It constitutes a diverse assemblage that includes a number of lizard-like forms, as well as more diverse types such as the pareiasaurs. Lee 1995, 1996, 1997 argues that turtles evolved from pareiasaurs, but this view is no longer considered likely. Rieppel and deBraga 1996 and deBraga and Rieppel, 1997 argue that turtles evolved from sauropterygians, and there is both molecular and fossil (''Pappochelys'') evidence for the origin of turtles among diapsid reptiles. Classification The foll ...
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Millerettidae
Millerettidae is an extinct family of parareptiles from the Middle Permian to the Late Permian period ( Capitanian - Changhsingian stages) of South Africa. The millerettids were small insectivores and probably resembled modern lizards in appearance and lifestyle. The following cladogram shows the phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ... position of the Millerettidae, from Ruta ''et al.'', 2011. References External linksBerkeley University information. Permian reptiles Prehistoric reptiles of Africa Guadalupian first appearances Lopingian extinctions Prehistoric reptile families {{Permian-reptile-stub ...
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