Orthoceratoidea
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Orthoceratoidea
Orthoceratoidea is a major subclass of nautiloid cephalopods. Members of this subclass usually have orthoconic (straight) to slightly cyrtoconic (curved) shells, and central to subcentral siphuncles which may bear internal deposits. Orthoceratoids are also characterized by dorsomyarian muscle scars (a small number of large scars concentrated at the top of the body chamber), extensive cameral deposits, and calciosiphonate connecting rings with a porous and calcitic inner layer. Currently, Orthoceratoidea comprises the orders Riocerida, Dissidocerida, Actinocerida, Pseudorthocerida, Lituitida and Orthocerida. Orthocerida is a noteworthy paraphyletic order which is ancestral to the major cephalopod groups such as the extinct ammonoids and living coleoids (cephalopods without external shells, including squids, octopus, cuttlefish, etc.). Taxonomy As a superorder, Orthoceratiodea was one of six superorders within the Nautiloidea, the others being the Plectronoceratoidea (= El ...
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Nautiloid
Nautiloids are a group of marine cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living ''Nautilus'' and '' Allonautilus''. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms (orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day. In a broad sense, "nautiloid" refers to a major cephalopod subclass or collection of subclasses (Nautiloidea ''sensu lato''). Nautiloids are typically considered one of three main groups of cephalopods, along with the extinct ammonoids (ammonites) and living coleoids (such as squid, octopus, and kin). While ammonoids and coleoids are monophyletic clades with exclusive ancestor-descendant re ...
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Orthoceratoidea
Orthoceratoidea is a major subclass of nautiloid cephalopods. Members of this subclass usually have orthoconic (straight) to slightly cyrtoconic (curved) shells, and central to subcentral siphuncles which may bear internal deposits. Orthoceratoids are also characterized by dorsomyarian muscle scars (a small number of large scars concentrated at the top of the body chamber), extensive cameral deposits, and calciosiphonate connecting rings with a porous and calcitic inner layer. Currently, Orthoceratoidea comprises the orders Riocerida, Dissidocerida, Actinocerida, Pseudorthocerida, Lituitida and Orthocerida. Orthocerida is a noteworthy paraphyletic order which is ancestral to the major cephalopod groups such as the extinct ammonoids and living coleoids (cephalopods without external shells, including squids, octopus, cuttlefish, etc.). Taxonomy As a superorder, Orthoceratiodea was one of six superorders within the Nautiloidea, the others being the Plectronoceratoidea (= El ...
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Cephalopod
A cephalopod is any member of the molluscan class Cephalopoda ( Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by '' Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been ide ...
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Pseudorthocerida
Pseudorthocerida is an order of generally straight longiconic orthoceratoids with a subcentral to marginal cyrtochoanitic siphuncle composed of variably expanded segments which may contain internal deposits that may develop into a continuous parietal lining. (Sweet 1964). Cameral deposits are common and concentrated ventrally. Apices typically have a slight to moderate exogastric curvature The Pseudorthocerida are included in a broad in-group of generally orthoconic cephalopods known as the Orthoceratoidea (Kroger 2008) along with the Ascocerida, Dissidocerida, Lituitida, and Orthocerida. The Pseudorthocerida were among the last living orthoconic nautiloids. One family, the Trematoceratidae, survived into the Triassic Period. Taxonomy Current understanding By current understanding the Pseudorthocerida contains the following families. : Pseudorthoceratidae : Cayutoceratidae : Pseudactinoceratidae : Spyroceratidae : Carbactinoceratidae : Trematoceratidae The Pseudorth ...
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Endocerida
Endocerida is an extinct nautiloid order, a group of cephalopods from the Lower Paleozoic with cone-like deposits in their siphuncle. Endocerida was a diverse group of cephalopods that lived from the Early Ordovician possibly to the Late Silurian. Their shells were variable in form. Some were straight (orthoconic), others curved (cyrtoconic); some were long (longiconic), others short (breviconic). Some long-shelled forms like '' Endoceras'' attained shell lengths close to . The related ''Cameroceras'' is anecdotally reported to have reached lengths approaching , but these claims are problematic. The overwhelming majority of endocerids and nautiloids in general are much smaller, usually less than a meter long when fully grown. Morphology Endocerids had a relatively small body chamber as well as a proportionally large siphuncle, which in some genera reached nearly half the shell diameter. This suggests that much of the visceral mass may have been housed within the siphuncle ...
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Troedssonellidae
Troedssonellidae is a family of orthoceroid cephalopods from the Ordovician, derived from rod-bearing Baltoceratidae, that have a continuous lining within the siphuncle that resembles very thin and slender endocones. Shells are generally slender and orthoconic. The siphuncle is central or subcentral, composed of straight or slightly expanded segments. Septal necks generally short and connecting rings are thin. Thin cameral deposits (lining the chambers) are known, which along with the position of the siphuncle and thin connecting rings distinguishes them from the endocerids in which they have been included.Flower 1976. Ordovician Cephalopod Faunas and Their Role in Correlation, in The Ordovician System: proceedings of a Palaeontological Association symposium, Sept 1974 Troedsonnellids first appear high in Lower Ordovician (Cassinian) strata, beginning with '' Tajaroceras'' and extend at least through the Whiterock Stage The Whiterockian, often referred to simply as the Whit ...
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Riocerida
Riocerida, originally named as Rioceratida, is an extinct order of Ordovician nautiloid cephalopods. They were the earliest-diverging members of the major group Orthoceratoidea, with which they share dorsomyarian muscle scars (a few large muscle scars concentrated at the top of the body chamber). However, they also possess several plesiomorphic ("primitive") traits which are absent in other orthoceratoids: Cameral and endosiphuncular deposits are absent, and the siphuncle has a marginal position along the lower edge of the shell. The conch (external shell) shape is slender, usually orthoconic (straight) or weakly cyrtoconic (slightly curved). The monophyly of Riocerida as a unique group is plausible but difficult to confirm, and they may instead be a paraphyletic grade or polyphyletic A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as ...
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Lituitida
Lituitida is an order of orthoceratoid cephalopods. They correspond to the family Lituitidae of the Treatise (Furnish & Glenister, 1964), reranked as an order and combined with other orthoceratoids. They are considered to be more closely related to the Orthocerida than to the Ascocerida or Pseudorthocerida Pseudorthocerida is an order of generally straight longiconic orthoceratoids with a subcentral to marginal cyrtochoanitic siphuncle composed of variably expanded segments which may contain internal deposits that may develop into a continuous p ... which are also included. Lituitids are characterized by smooth to annulate shells in which the juvenile portion near the apex is coiled or cyrtoconic. The adult portion is straight or slightly sigmoidal and may be expanded. Growth lines show a deep hyponomic sinus and lateral ocular sinuses. The siphuncle is subcentral with layered connecting rings. Cameral deposits are prevalent and concentrated more ventrally. The apex is s ...
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Orthocone
An orthocone is an unusually long straight shell of a nautiloid cephalopod.; During the 18th and 19th centuries, all shells of this type were named '' Orthoceras'', creating a wastebasket taxon, but it is now known that many groups of nautiloids developed or retained this type of shell. An orthocone can be thought of as like a '' nautilus'', but with the shell straight and uncoiled. It was previously believed that these represented the most primitive form of nautiloid, but it is now known that the earliest nautiloids had shells that were slightly curved. An orthoconic form evolved several times among cephalopods, and among nautiloid cephalopods is prevalent among the ellesmerocerids, endocerids, actinocerids, orthoceratoids, and bactritids. Orthocones existed from the Late Cambrian to the Late Triassic, but they were most common in the early Paleozoic. Revivals of the orthocone design later occurred in other cephalopod groups, notably baculitid ammonites in the Cretaceo ...
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Plectronoceratoidea
Plectronoceratoidea is a superorder or subclass containing primitive nautiloids from the Late Cambrian and Early Ordovician. This group is best considered a paraphyletic grade of early cephalopods, as it contains the ancestors of subsequent post-Cambrian cephalopod orders. Plectronoceratoidea contains several exclusively Cambrian cephalopod orders: Plectronocerida, Protactinocerida, and Yanhecerida. Under some classification schemes, the plectronoceratoid grade may also contain the paraphyletic Cambro-Ordovician Ellesmerocerida. A few older studies consider Plectronocerida to be descended from Ellesmerocerida (rather than ancestral), but this is no longer believed to be the case. The Plectronocerida is the earliest group of the four, and may have given rise to the other three. Of these four orders, only the Ellesmerocerida crossed (barely) into the Ordovician, with two genera, ''Ectenolites ''Ectenolites'' is a genus of small, slender, cylindrical Ellesmeroceratids that res ...
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Orthocerida
Orthocerida is an order of extinct Orthoceratoid cephalopods also known as the Michelinocerida that lived from the Early Ordovician () possibly to the Late Triassic (). A fossil found in the Caucasus suggests they may even have survived until the Early Cretaceous (). They were most common however from the Ordovician to the Devonian. Shell form The shell is usually long, and may be straight (" orthoconic") or gently curved. In life, these animals may have been similar to the modern squid, except for the long shell. The internal structure of the shell consists of concavo-convex chambers linked by a centrally-placed tube called a siphuncle. There is a tendency for the chambers to develop cameral deposits, which were used as ballast to balance the long gas-filled shell. Depending on the family, the siphuncle has orthochoanitic (short and straight) or cyrtochoanitic (outwardly curved) septal necks, which protrude from the septa. The shell surface may be (depending on the spe ...
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Dissidocerida
Dissidocerida is an order of Early Ordovician to the Early Silurian orthoceratoid cephalopods in which the siphuncle has a continuous lining or a longitudinal rod-like structure within. The order Dissidocerida was proposed by Zhuravleva (1994) for ''Dissidoceras undosum'' Zhuravleva, 1964, a Silurian orthoconic genus with a wide, nearly central, tubular siphuncle, suborthochoanitic septal necks, and endosiphuncular rod; in many aspects similar to rod-bearing orthocones of the Ordovician. It is thought by some that Ordovician-Silurian orthocones with continuous longitudinal endosiphuncular laminar deposits constitute a single phylogenetic stock. The Dissidocerida as understood includes the Polymeridae, Rangeroceratidae, Troedssonellidae, and Dissidocertidae. The Polymeridae and Rangeroceratidae were proposed by Evans in 2005 for ''Polymeres'' Murchison 1839 and ''Rangeroceras'' Hook and Flower 1977, respectively. Troedssonellidae Troedssonellidae is a family of orthoce ...
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