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Opabiniidae
Opabiniidae is an extinct family of bizarre marine stem-arthropods. Its type and best-known genus is '' Opabinia''. It also contains '' Utaurora''. Opabiniids closely resemble radiodonts, but their frontal appendages were basally fused into a proboscis. Opabiniids also distinguishable from radiodonts by setal blades covering at least part of the body flaps and serrated caudal rami. History of study Opabiniidae was named by Charles Doolittle Walcott in 1912, alongside its type species ''Opabinia''. Walcott interpreted Opabiniidae as a family of anostracan crustaceans, most closely related to Thamnocephalidae. ''Opabinia'' was restudied in the 1970s, and reinterpreted as a stranger animal. Stephen Jay Gould referred to ''Opabinia'' as a "weird wonder", and an illustration of ''Opabinia'' prompted laughter when it was first revealed at a paleontological conference. In 2022, a second opabiniid, '' Utaurora'', was identified. '' Myoscolex'' from Emu Bay Shale is sometimes sugge ...
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Radiodonta
Radiodonta is an extinct order of stem-group arthropods that was successful worldwide during the Cambrian period. They may be referred to as radiodonts, radiodontans, radiodontids, anomalocarids, or anomalocaridids, although the last two originally refer to the family Anomalocarididae, which previously included all species of this order but is now restricted to only a few species. Radiodonts are distinguished by their distinctive frontal appendages, which are morphologically diverse and used for a variety of functions. Radiodonts included the earliest large predators known, but they also included sediment sifters and filter feeders. Some of the most famous species of radiodonts are the Cambrian taxa ''Anomalocaris canadensis'', ''Hurdia victoria'', '' Peytoia nathorsti'', '' Titanokorys gainessii, Cambroraster falcatus'' and '' Amplectobelua symbrachiata'', the Ordovician '' Aegirocassis benmoulai'' and the Devonian ''Schinderhannes bartelsi''. Etymology The name Radiodonta (Lat ...
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Mieridduryn
''Mieridduryn'' is a genus of extinct dinocaridid arthropod that lived during the Middle Ordovician of what is now the United Kingdom. This animal was described in 2022 based on a singular fossil found in Castle Bank, a Burgess shale type lagerstätte located in the country of Wales. This animal's taxonomic affinities are somewhat unclear, but there are some hypotheses. One is that this animal represents a new grade of Stem- euarthropods that evolved features similar to the Cambrian aged opabiniids (a family of basal arthropods). Another is that if the features seen in ''Mieridduryn'' are convergent, and not homologous, to those seen in radiodonts, then this animal represents a late surviving opabiniid (if the latter option is correct, then this means the opabiniids survived for over 40 million years longer than what was once thought). Free abstract at History of research and etymology The one specimen of this species was found in Castle Bank, and was described in 202 ...
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Utaurora
''Utaurora'' is an extinct genus of opabiniid, which were bizarre stem-arthropods closely related to true arthropods and radiodonts; the type species is ''U. comosa''. The animal's fossils come from the Cambrian of Utah. This genus is so far the only other known unquestionable opabiniid, with the other being ''Opabinia'' itself. There are other animals like '' Myoscolex'' and ''Mieridduryn'' that could be opabiniids, but the classification of those two genera is still debated. History of study The holotype specimen of ''Utaurora comosa'', KUMIP 314087, was collected from the Wheeler Formation in Utah. It was initially described as a specimen of ''Anomalocaris'' in 2008. In 2022, Pates ''et al.'' reinterpreted the specimen as an opabiniid and described it as a new genus and species. Etymology ''Utaurora'' is a portmanteau of Utah, in reference to where the specimen was found, and Aurora, the name of a Roman goddess. The reference to Aurora was chosen as she is a goddess of the ...
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Arthropods
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior organs; it has an open circulatory system. Like their exteriors, the internal organs of arthropods are generally built of repeated segments. Their nervous system is " ...
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Myoscolex
''Myoscolex'' is an early animal species known from the Cambrian Emu Bay Shale in South Australia. It was interpreted as an annelid and some supports that theory, while it was also considered as an arthropod close to '' Opabinia'', through other studies questions that interpretation. ''Myoscolex'' is the earliest known example of phosphotized muscle tissue, and as to which shows distinct annulation. Etymology ''Myoscolex ateles'' derives its name from Greek, "Myo-" meaning muscle, "Scolex" meaning worm, and "Ateles" meaning incomplete. This definition of "Incomplete muscle worm" is due to its initial identification as a "muscular" annelid worm with indistinct features after fossilization. Description As an Opabiniid When described as an Opabiniid, ''Myoscolex'' is said to have at least 3 eyes, a thin proboscis jutting from under the proposed eyes, lateral lobes on the trunk, a tail fan on the posterior segments, and an upwardly curving trunk. It would have lived as a fast ...
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Middle Cambrian
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Crustaceans
Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group can be treated as a subphylum under the clade Mandibulata. It is now well accepted that the hexapods emerged deep in the Crustacean group, with the completed group referred to as Pancrustacea. Some crustaceans (Remipedia, Cephalocarida, Branchiopoda) are more closely related to insects and the other hexapods than they are to certain other crustaceans. The 67,000 described species range in size from '' Stygotantulus stocki'' at , to the Japanese spider crab with a leg span of up to and a mass of . Like other arthropods, crustaceans have an exoskeleton, which they moult to grow. They are distinguished from other groups of arthropods, such as insects, myriapods and chelicerates, by the possession of biramous (two-parted) limbs, and by ...
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Middle Ordovician
The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. The Ordovician, named after the Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official period of the Paleozoic Era by the International Geological Congress. Life continued to flourish during the Ordovician as it did in the earlier Cambrian Pe ...
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Panarthropoda
Panarthropoda is a proposed animal clade containing the extant phyla Arthropoda, Tardigrada (water bears) and Onychophora (velvet worms). Panarthropods also include extinct marine legged worms known as lobopodians ("Lobopodia"), a paraphyletic group where the last common ancestor and basal members (stem-group) of each extant panarthropod phylum are thought to have risen. However the term "Lobopodia" is sometimes expanded to include tardigrades and onychophorans as well. Common characteristics of the Panarthropoda include a segmented body, paired ladder-like, ventral nervous system, and the presence of paired appendages correlated with body segments. Taxonomy Not all studies support the monophyly of Panarthropoda, but most do, including neuroanatomical, phylogenomic and palaeontological studies. At least a close relationship between onychophorans and arthropods is widely agreed upon, but the position of tardigrades is more controversial. Some phylogenomic studies h ...
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Emu Bay Shale
The Emu Bay Shale is a geological formation in Emu Bay, South Australia, containing a major Konservat-Lagerstätte (fossil beds with soft tissue preservation). It is one of two in the world containing Redlichiidan trilobites. The Emu Bay Shale is dated as Cambrian Series 2, Stage 4, correlated with the upper Botomian Stage of the Lower Cambrian. Its mode of preservation is the same as the Burgess shale, but the larger grain size of the Emu Bay rock means that the quality of preservation is lower. More than 50 species of trilobites, non-biomineralized arthropods, palaeoscolecids, a lobopodian, a polychaete, vetulicolians, nectocaridids, hyoliths, brachiopods, sponges, chancelloriids, and a chelicerate are known from the Emu Bay Shale. Description The Emu Bay Shale of Kangaroo Island, South Australia, is Australia's only known Burgess-Shale-type Konservat-Lagerstätte, and includes faunal elements such as ''Anomalocaris'', ''Tuzoia'', ''Isoxys'', and '' Wronascolex'', i ...
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Thamnocephalidae
Thamnocephalidae is a family of crustaceans with wide distribution including Western Australia, Southern Africa and South America. It was originally described as a subfamily of Branchipodidae by Alpheus Spring Packard in 1883, and elevated to the rank of family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ... by Simon in 1886. Six genera are recognised, in two subfamilies: ;Thamnocephalinae *'' Thamnocephalus'' Packard, 1879 *'' Carinophallus'' Rogers, 2006 ;Branchinellinae *'' Dendrocephalus'' Daday, 1908 *'' Phallocryptus'' Birabén, 1951 *'' Spiralifrons'' Dixon, 2010 *'' Branchinella'' Sayce, 1903 References Anostraca Crustacean families Taxa named by Alpheus Spring Packard {{branchiopoda-stub ...
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20220214 Opabiniidae Size
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