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Orovenator Mayorum
''Orovenator'' is an extinct genus of diapsid from Lower Permian (Artinskian stage) deposits of Oklahoma, United States. It is known from two partial skulls from the Richards Spur locality in Oklahoma. The holotype OMNH 74606 consists of a partial skull preserving snout and mandible, and the referred specimen, OMNH 74607, a partial skull preserving the skull roof, vertebrae and palatal elements. It was first named by Robert R. Reisz, Sean P. Modesto and Diane M. Scott in 2011 and the type species is ''Orovenator mayorum''. The generic name means "mountain", ''oro'', in Greek in reference to the Richards Spur locality, which was mountainous during the Permian period and "hunter", ''venator'', in Latin. The specific name honours Bill and Julie May. ''Orovenator'' is the oldest and most basal neodiapsid to date. A 2018 redescription by David Ford and Roger Benson found that ''Orovenator'' shared many similarities with varanopids, a group of reptile-like tetrapods tradit ...
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Artinskian
In the geologic timescale, the Artinskian is an age (geology), age or stage (stratigraphy), stage of the Permian. It is a subdivision of the Cisuralian Epoch (geology), Epoch or series (stratigraphy), Series. The Artinskian likely lasted between and megaannum, million years ago (Ma) according to the most recent revision of the International Commission on Stratigraphy (ICS) in 2022. It was preceded by the Sakmarian and followed by the Kungurian. Stratigraphy The Artinskian is named after the goniatite grits of Artinsk which was introduced by Roderick Murchison, Édouard de Verneuil and count Alexander von Keyserling in their ''The Geology of Russia in Europe and the Ural Mountains'' (1845). The grits of Artinsk, in turn, get its name from the Artinsky District with center in the Russian smalltown of Arti, Russia, Arti (formerly ''Artinsk zavod''), situated in the middle Ural Mountains, Urals, about 170 km southwest of Yekaterinburg. The stage was introduced into scientific li ...
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Greek (language)
Greek (, ; , ) is an Indo-European languages, Indo-European language, constituting an independent Hellenic languages, Hellenic branch within the Indo-European language family. It is native to Greece, Cyprus, Italy (in Calabria and Salento), southern Albania, and other regions of the Balkans, Caucasus, the Black Sea coast, Asia Minor, and the Eastern Mediterranean. It has the list of languages by first written accounts, longest documented history of any Indo-European language, spanning at least 3,400 years of written records. Its writing system is the Greek alphabet, which has been used for approximately 2,800 years; previously, Greek was recorded in writing systems such as Linear B and the Cypriot syllabary. The Greek language holds a very important place in the history of the Western world. Beginning with the epics of Homer, ancient Greek literature includes many works of lasting importance in the European canon. Greek is also the language in which many of the foundational texts ...
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Cladogram
A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other e ...
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Heleosaurus
''Heleosaurus scholtzi'' is an extinct species of basal synapsids, known as pelycosaurs, in the family of Varanopidae during the middle Permian.Reisz, R. R. & Modesto, S. P. 2007. ''Heleosaurus scholtzi'' from the Permian of South Africa: a varanopid synapsid, not a diapsid reptile. ''Journal of Vertebrate Paleontology'' 27 (3): 734-739. At first ''H. scholtzi'' was mistakenly classified as a diapsid. Members of this family were carnivorous and had dermal armor, and somewhat resembled monitor lizards. This family was the most geologically long lived, widespread, and diverse group of early amniotes. To date only two fossils have been found in the rocks of South Africa.Broom, R. "On Some New Fossil Reptiles from the Karroo Beds of Victoria West, South Africa." ''Transactions of the South African Philosophical Society'' 18 (1907): 31-42. Taylor and Francis Online. Web. Botha-Brink, Jennifer. "A Mixed-Age Classed 'Pelycosaur' Aggregation from South Africa: Earliest Evidence of Parental ...
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Mammal
A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three Evolution of mammalian auditory ossicles, middle ear bones. These characteristics distinguish them from reptiles and birds, from which their ancestors Genetic divergence, diverged in the Carboniferous Period over 300 million years ago. Around 6,640 Neontology#Extant taxon, extant species of mammals have been described and divided into 27 Order (biology), orders. The study of mammals is called mammalogy. The largest orders of mammals, by number of species, are the rodents, bats, and eulipotyphlans (including hedgehogs, Mole (animal), moles and shrews). The next three are the primates (including humans, monkeys and lemurs), the Artiodactyl, even-toed ungulates (including pigs, camels, and whales), and the Carnivora (including Felidae, ...
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Amniote
Amniotes are tetrapod vertebrate animals belonging to the clade Amniota, a large group that comprises the vast majority of living terrestrial animal, terrestrial and semiaquatic vertebrates. Amniotes evolution, evolved from amphibious Stem tetrapoda, stem tetrapod ancestors during the Carboniferous geologic period, period. Amniota is defined as the smallest crown clade containing humans, the Greek tortoise, and the Nile crocodile. Amniotes are distinguished from the other living tetrapod clade — the anamniote, non-amniote lissamphibians (frogs/toads, salamanders/newts and caecilians) — by: the development of three fetal membranes, extraembryonic membranes (amnion for embryonic protection, chorion for gas exchange, and allantois for metabolic waste disposal or storage); thicker and keratinized skin; rib, costal respiration (breathing by expanding/constricting the rib cage); the presence of adrenal cortex, adrenocortical and chromaffin cell, chromaffin tissues as adrenal g ...
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Synapsid
Synapsida is a diverse group of tetrapod vertebrates that includes all mammals and their extinct relatives. It is one of the two major clades of the group Amniota, the other being the more diverse group Sauropsida (which includes all extant reptiles and therefore, birds). Unlike other amniotes, synapsids have a single temporal fenestra, an opening low in the skull roof behind each eye socket, leaving a zygomatic arch, bony arch beneath each; this accounts for the name "synapsid". The distinctive temporal fenestra developed about 318 million years ago during the Late Carboniferous period, when synapsids and sauropsids diverged, but was subsequently merged with the orbit in early mammals. The basal (phylogenetics), basal amniotes (reptiliomorphs) from which synapsids evolved were historically simply called "reptiles". Therefore, stem group synapsids were then described as mammal-like reptiles in classical systematics, and non-therapsid synapsids were also referred to as pelyco ...
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Tetrapod
A tetrapod (; from Ancient Greek :wiktionary:τετρα-#Ancient Greek, τετρα- ''(tetra-)'' 'four' and :wiktionary:πούς#Ancient Greek, πούς ''(poús)'' 'foot') is any four-Limb (anatomy), limbed vertebrate animal of the clade Tetrapoda (). Tetrapods include all Neontology#Extant taxa versus extinct taxa, extant and Extinction, extinct amphibians and amniotes, with the latter in turn Evolution, evolving into two major clades, the Sauropsida, sauropsids (reptiles, including dinosaurs and therefore birds) and synapsids (extinct pelycosaur, "pelycosaurs", therapsids and all extant mammals, including Homo sapiens, humans). Hox gene mutations have resulted in some tetrapods becoming Limbless vertebrate, limbless (snakes, legless lizards, and caecilians) or two-limbed (cetaceans, sirenians, Bipedidae, some lizards, kiwi (bird), kiwis, and the extinct moa and elephant birds). Nevertheless, they still qualify as tetrapods through their ancestry, and some retain a pair of ves ...
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Varanopidae
Varanopidae is an extinct family (biology), family of amniotes known from the Late Carboniferous to Middle Permian that resembled monitor lizards (with the name of the group deriving from the monitor lizard genus ''Varanus'') and may have filled a similar niche. Typically, they are considered to be relatively Basal (phylogenetics), basal Synapsida, synapsids (and thus more closely related to mammals than to reptiles), although some studies from the late 2010s recovered them being taxonomically closer to diapsid reptiles, recent studies from the early 2020s support their traditional placement as synapsids on the basis of high degree of bone labyrinth ossification, maxillary canal morphology and phylogenetic analyses. A varanopid from the late Middle Permian Pristerognathus Assemblage Zone, ''Pristerognathus'' Assemblage Zone (Capitanian) is the youngest known varanopid and the last member of the "pelycosaur" group of synapsids. Thus, Varanopidae vanishes from the fossil record at th ...
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Roger B
Roger is a masculine given name, and a surname. The given name is derived from the Old French personal names ' and '. These names are of Germanic languages">Germanic origin, derived from the elements ', ''χrōþi'' ("fame", "renown", "honour") and ', ' ("spear", "lance") (Hrōþigēraz). The name was introduced into England by the Normans. In Normandy, the Franks, Frankish name had been reinforced by the Old Norse cognate '. The name introduced into England replaced the Old English cognate '. ''Roger'' became a very common given name during the Middle Ages. A variant form of the given name ''Roger'' that is closer to the name's origin is '' Rodger''. Slang and other uses From up to , Roger was slang for the word "penis". In ''Under Milk Wood'', Dylan Thomas writes "jolly, rodgered" suggesting both the sexual double entendre and the pirate term "Jolly Roger". In 19th-century England, Roger was slang for another term, the cloud of toxic green gas that swept through the chlori ...
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Orovenator
''Orovenator'' is an extinct genus of diapsid from Lower Permian ( Artinskian stage) deposits of Oklahoma, United States. It is known from two partial skulls from the Richards Spur locality in Oklahoma. The holotype OMNH 74606 consists of a partial skull preserving snout and mandible, and the referred specimen, OMNH 74607, a partial skull preserving the skull roof, vertebrae and palatal elements. It was first named by Robert R. Reisz, Sean P. Modesto and Diane M. Scott in 2011 and the type species is ''Orovenator mayorum''. The generic name means "mountain", ''oro'', in Greek in reference to the Richards Spur locality, which was mountainous during the Permian period and "hunter", ''venator'', in Latin. The specific name honours Bill and Julie May. ''Orovenator'' is the oldest and most basal neodiapsid to date. A 2018 redescription by David Ford and Roger Benson found that ''Orovenator'' shared many similarities with varanopids, a group of reptile-like tetrapo ...
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Neodiapsid
Diapsids ("two arches") are a clade of sauropsids, distinguished from more primitive eureptiles by the presence of two holes, known as temporal fenestrae, in each side of their skulls. The earliest traditionally identified diapsids, the Araeoscelidia, araeoscelidians, appeared about three hundred million years ago during the late Carboniferous period. All diapsids other than the most primitive ones in the clade Araeoscelidia are often placed into the clade Neodiapsida. The diapsids are extremely diverse, and include birds and all modern reptile groups, including turtles, which were historically thought to lie outside the group. All modern reptiles and birds are placed within the neodiapsid subclade Sauria. Although some diapsids have lost either one hole (lizards), or both holes (snakes and turtles), or have a heavily restructured skull (modern birds), they are still scientific classification, classified as diapsids based on their ancestry. At least 17,084 species of diapsid animals ...
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