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Nyasasaurus
''Nyasasaurus'' (meaning " Lake Nyasa lizard") is an extinct genus of avemetatarsalian archosaur from the putatively Middle Triassic Manda Formation of Tanzania that may be the earliest known dinosaur. The type species ''Nyasasaurus parringtoni'' was first described in 1956 in the doctoral thesis of English paleontologist Alan J. Charig, but it was not formally described until 2013. Previously, the oldest record of dinosaurs was from Brazil and Argentina and dated back to the mid-late Carnian stage, about 233.23 to 231.4 million years ago. ''Nyasasaurus'' comes from a deposit formerly considered Anisian in age, meaning that it would predate other early dinosaurs by about 12 million years. However, more recent studies cast doubt to this age, pointing out that the deposits would actually be Carnian in age, which would considerably reduce this temporal gap. History of study In the 1930s, the holotype of ''Nyasasaurus'' was collected in Parrington's locality B36 from the Lifua Memb ...
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Nyasasaurus
''Nyasasaurus'' (meaning " Lake Nyasa lizard") is an extinct genus of avemetatarsalian archosaur from the putatively Middle Triassic Manda Formation of Tanzania that may be the earliest known dinosaur. The type species ''Nyasasaurus parringtoni'' was first described in 1956 in the doctoral thesis of English paleontologist Alan J. Charig, but it was not formally described until 2013. Previously, the oldest record of dinosaurs was from Brazil and Argentina and dated back to the mid-late Carnian stage, about 233.23 to 231.4 million years ago. ''Nyasasaurus'' comes from a deposit formerly considered Anisian in age, meaning that it would predate other early dinosaurs by about 12 million years. However, more recent studies cast doubt to this age, pointing out that the deposits would actually be Carnian in age, which would considerably reduce this temporal gap. History of study In the 1930s, the holotype of ''Nyasasaurus'' was collected in Parrington's locality B36 from the Lifua Memb ...
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Manda Formation
The Manda Formation (also known as the Manda Beds) is a Middle Triassic (Anisian?) or possibly Late Triassic (Carnian?) geologic formation in Tanzania. It preserves fossils of many terrestrial vertebrates from the Triassic, including some of the earliest dinosauromorph archosaurs. The formation is often considered to be Anisian in age according to general tetrapod biochronology hypotheses and correlations to the ''Cynognathus'' Assemblage Zone of South Africa. However, some recent studies cast doubt to this age, suggesting that parts deposits may actually be younger (Carnian) in age. History of study One of the first to study rocks of the Manda Formation was British geologist G. M. Stockley. In 1932, Stockley explored the geology of the Ruhuhu Basin in Tanzania. He called a series of layers dating from the Late Carboniferous to the Middle Triassic the Songea Series and divided it into eight units labelled K1-K8. Stockley was also the first to describe fossils from these rocks, n ...
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Francis Rex Parrington
Francis Rex Parrington (20 February 1905 – 17 April 1981) was a British vertebrate palaeontologist and comparative anatomist at the University of Cambridge. A Fellow of the Royal Society, he was director of the Cambridge University Museum of Zoology and past president of the zoology section of the British Association for the Advancement of Science. Early life Francis Rex Parrington was born on 20 February 1905 in Bromborough near Neston in Cheshire, the youngest of three children of brewery owner Frank Harding Parrington and his wife Bessie Harding. After the death of his father in 1907, his mother remarried and 'Rex' Parrington was brought up in Central Liverpool, where his stepfather was a general practitioner. His interest in natural history developed in childhood, and he collected wildflowers, beetles, and fossils. In 1920, his family moved to North Wales, where he could freely explore his interests in the natural world. He was educated at the Merchant Taylors' School in ...
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Sterling Nesbitt
Sterling Nesbitt (born March 25, 1982, in Mesa, Arizona) is an American paleontologist best known for his work on the origin and early evolutionary patterns of archosaurs. He is currently an associate professor at Virginia Tech in the Department of Geosciences. Biography Sterling Nesbitt received his B.A. in integrative biology with a minor in geology from the University of California Berkeley in 2004. He received his PhD from Columbia University in 2009, completing the majority of his research at the American Museum of Natural History in New York City. He subsequently held postdoctoral researcher positions at the University of Texas at Austin, the University of Washington, and the Field Museum. He is currently an associate professor in thDepartment of Geosciencesat Virginia Tech in Blacksburg, Virginia. He is also a research associate/affiliate of the American Museum of Natural History, the Vertebrate Paleontology Lab at The University of Texas at Austin, the Virginia Museum of N ...
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Dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is the subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya; their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic, morphological and ecological standpoints. Birds, at over 10,700 living species, ar ...
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Avemetatarsalia
Avemetatarsalia (meaning "bird metatarsals") is a clade of diapsid reptiles containing all archosaurs more closely related to birds than to crocodilians. The two most successful groups of avemetatarsalians were the dinosaurs and pterosaurs. Dinosaurs were the largest terrestrial animals for much of the Mesozoic Era, and one group of small feathered dinosaurs (Aves, i.e. birds) has survived up to the present day. Pterosaurs were the first flying vertebrates and persisted through the Mesozoic before dying out at the Cretaceous-Paleogene (K-Pg) extinction event. Both dinosaurs and pterosaurs appeared in the Triassic Period, shortly after avemetatarsalians as a whole. The name Avemetatarsalia was first established by British palaeontologist Michael Benton in 1999. An alternate name is Panaves, or "all birds", in reference to its definition containing all animals, living or extinct, which are more closely related to birds than to crocodilians. Although dinosaurs and pterosaurs w ...
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Archosaur
Archosauria () is a clade of diapsids, with birds and crocodilians as the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term which includes birds. Extinct archosaurs include non-avian dinosaurs, pterosaurs, and extinct relatives of crocodilians. Modern paleontologists define Archosauria as a crown group that includes the most recent common ancestor of living birds and crocodilians, and all of its descendants. The base of Archosauria splits into two clades: Pseudosuchia, which includes crocodilians and their extinct relatives, and Avemetatarsalia, which includes birds and their extinct relatives (such as non-avian dinosaurs and pterosaurs). Older definitions of the group Archosauria rely on shared morphological characteristics, such as an antorbital fenestra in the skull, serrated teeth, and an upright stance. Some extinct reptiles, such as proterosuchids and euparkeriids, possessed these features yet originate ...
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Middle Triassic
In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between Ma and Ma (million years ago). It is preceded by the Early Triassic Epoch and followed by the Late Triassic Epoch. The Middle Triassic is divided into the Anisian and Ladinian ages or stages. Formerly the middle series in the Triassic was also known as Muschelkalk. This name is now only used for a specific unit of rock strata with approximately Middle Triassic age, found in western Europe. Middle Triassic fauna Following the Permian–Triassic extinction event, the most devastating of all mass-extinctions, life recovered slowly. In the Middle Triassic, many groups of organisms reached higher diversity again, such as the marine reptiles (e.g. ichthyosaurs, sauropterygians, thallatosaurs), ray-finned fish and many invertebrate gr ...
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Thecodontosaurus
''Thecodontosaurus'' ("socket-tooth lizard") is a genus of herbivorous basal sauropodomorph dinosaur that lived during the late Triassic period ( Rhaetian age). Its remains are known mostly from Triassic "fissure fillings" in South England. ''Thecodontosaurus'' was a small bipedal animal, about 2 m (6.5 ft) long. It is one of the first dinosaurs to be discovered and is one of the oldest that existed. Many species have been named in the genus, but only the type species ''Thecodontosaurus antiquus'' is seen as valid today. Discovery and naming ''Thecodontosaurus antiquus'' In the autumn of 1834, surgeon Henry Riley (1797–1848) and the curator of the Bristol Institution, Samuel Stutchbury, began to excavate "saurian remains" at the quarry of Durdham Down, at Clifton, presently a part of Bristol, which is part of the Magnesian Conglomerate. In 1834 and 1835, they briefly reported on the finds. They provided their initial description in 1836, naming a new genus: ''T ...
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Carbonate
A carbonate is a salt of carbonic acid (H2CO3), characterized by the presence of the carbonate ion, a polyatomic ion with the formula . The word ''carbonate'' may also refer to a carbonate ester, an organic compound containing the carbonate group C(=O)(O–)2. The term is also used as a verb, to describe carbonation: the process of raising the concentrations of carbonate and bicarbonate ions in water to produce carbonated water and other carbonated beverageseither by the addition of carbon dioxide gas under pressure or by dissolving carbonate or bicarbonate salts into the water. In geology and mineralogy, the term "carbonate" can refer both to carbonate minerals and carbonate rock (which is made of chiefly carbonate minerals), and both are dominated by the carbonate ion, . Carbonate minerals are extremely varied and ubiquitous in chemically precipitated sedimentary rock. The most common are calcite or calcium carbonate, CaCO3, the chief constituent of limestone (as we ...
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Lungfish
Lungfish are freshwater vertebrates belonging to the order Dipnoi. Lungfish are best known for retaining ancestral characteristics within the Osteichthyes, including the ability to breathe air, and ancestral structures within Sarcopterygii, including the presence of lobed fins with a well-developed internal skeleton. Lungfish represent the closest living relatives of the tetrapods. Today there are only six known species of lungfish, living in Africa, South America, and Australia. The fossil record shows that lungfish were abundant since the Triassic. While vicariance would suggest this represents an ancient distribution limited to the Mesozoic supercontinent Gondwana, the fossil record suggests advanced lungfish had a widespread freshwater distribution and the current distribution of modern lungfish species reflects extinction of many lineages subsequent to the breakup of Pangaea, Gondwana and Laurasia. Lungfish have historically been referred to as salamanderfish, but ...
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Amphibia
Amphibians are four-limbed and ectothermic vertebrates of the class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terrestrial, fossorial, arboreal or freshwater aquatic ecosystems. Thus amphibians typically start out as larvae living in water, but some species have developed behavioural adaptations to bypass this. The young generally undergo metamorphosis from larva with gills to an adult air-breathing form with lungs. Amphibians use their skin as a secondary respiratory surface and some small terrestrial salamanders and frogs lack lungs and rely entirely on their skin. They are superficially similar to reptiles like lizards but, along with mammals and birds, reptiles are amniotes and do not require water bodies in which to breed. With their complex reproductive needs and permeable skins, amphibians are often ecological indicators; in recent decades there has been a dramati ...
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