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Novialoidea
Novialoidea (meaning "new wings") is an extinct clade of macronychopteran pterosaurs that lived from the latest Early Jurassic to the latest Late Cretaceous (early Toarcian to late Maastrichtian ageBarrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). Pterosaur distribution in time and space: an atlas. ''Zitteliana'', 61-107/ref>), their fossils having been found on all continents except Antarctica. History and classification Novialoidea was named by paleontologist Alexander Wilhelm Armin Kellner in 2003 as a node-based taxon consisting of the last common ancestor of '' Campylognathoides'', ''Quetzalcoatlus'' and all its descendants. This name was derived from Latin ''novus'' "new", and ''ala'', "wing", in reference to the wing synapomorphies that the members of the clade possess.Kellner, A. W. A., (2003): Pterosaur phylogeny and comments on the evolutionary history of the group. pp. 105-137. — ''in'' Buffetaut, E. & Mazin, J.-M., (eds.): ''Evolution and P ...
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Campylognathoides
''Campylognathoides'' ("curved jaw", Strand 1928) is an extinct genus of pterosaur discovered in the Württemberg Lias deposits (dated to the early Toarcian ageBarrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). "Pterosaur distribution in time and space: an atlas". ''Zitteliana'', 61–107/ref>) of Germany; this first specimen however, consisted only of wing fragments. Further better preserved specimens were found in the Holzmaden shale; based on these specimens, Felix Plieninger erected a new genus. Discovery In 1858 Friedrich August Quenstedt named a new species of ''Pterodactylus'': ''P. liasicus''. It was based on a fossil, holotype GPIT 9533, consisting of some wing bones, found on the Wittberg near Metzingen in layers dating from the early Toarcian, about 180 million years old. The specific name referred to the Lias. Quenstedt thought he had identified long metacarpals in the wing, concluding that the new species was therefore not belonging to more b ...
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Campylognathoides Liasicus
''Campylognathoides'' ("curved jaw", Strand 1928) is an extinct genus of pterosaur discovered in the Württemberg Lias deposits (dated to the early Toarcian ageBarrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). "Pterosaur distribution in time and space: an atlas". ''Zitteliana'', 61–107/ref>) of Germany; this first specimen however, consisted only of wing fragments. Further better preserved specimens were found in the Holzmaden shale; based on these specimens, Felix Plieninger erected a new genus. Discovery In 1858 Friedrich August Quenstedt named a new species of ''Pterodactylus'': ''P. liasicus''. It was based on a fossil, holotype GPIT 9533, consisting of some wing bones, found on the Wittberg near Metzingen in layers dating from the early Toarcian, about 180 million years old. The specific name referred to the Lias. Quenstedt thought he had identified long metacarpals in the wing, concluding that the new species was therefore not belonging to more b ...
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Pterosaur
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were i ...
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Macronychoptera
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insectiv ...
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Eudimorphodon Ranzii
''Eudimorphodon'' was a pterosaur that was discovered in 1973 by Mario Pandolfi in the town of Cene, Italy and described the same year by Rocco Zambelli. The nearly complete skeleton was retrieved from shale deposited during the Late Triassic (mid to late Norian stage),Wellnhofer, P. (1991). "Summary of Triassic Pterosaurs." ''The Illustrated Encyclopedia of Pterosaurs.'' London, UK: Salamander Books Limited. p. 67. . making ''Eudimorphodon'' one of the oldest pterosaurs known.Cranfield, I. ''The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures''. London: Salamander Books, Ltd. Pp. 280–281. It had a wingspan of about and at the end of its long bony tail may have been a diamond-shaped flap like in the later ''Rhamphorhynchus''. If so, the flap may have helped it steer while maneuvering in the air. ''Eudimorphodon'' is known from several skeletons, including juvenile specimens. Discovery and species ''Eudimorphodon'' currently includes one species, the type s ...
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Rhamphorhynchus
''Rhamphorhynchus'' (, from Ancient Greek ''rhamphos'' meaning "beak" and ''rhynchus'' meaning "snout") is a genus of long-tailed pterosaurs in the Jurassic period. Less specialized than contemporary, short-tailed pterodactyloid pterosaurs such as ''Pterodactylus'', it had a long tail, stiffened with ligaments, which ended in a characteristic soft-tissue tail vane. The mouth of ''Rhamphorhynchus'' housed needle-like teeth, which were angled forward, with a curved, sharp, beak-like tip lacking teeth, indicating a diet mainly of fish; indeed, fish and cephalopod remains are frequently found in ''Rhamphorhynchus'' abdominal contents, as well as in their coprolites. Although fragmentary fossil remains possibly belonging to ''Rhamphorhynchus'' have been found in England, Tanzania, and Spain, the best preserved specimens come from the Solnhofen limestone of Bavaria, Germany. Many of these fossils preserve not only the bones but impressions of soft tissues, such as wing membranes. Scat ...
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Eudimorphodon
''Eudimorphodon'' was a pterosaur that was discovered in 1973 by Mario Pandolfi in the town of Cene, Italy and described the same year by Rocco Zambelli. The nearly complete skeleton was retrieved from shale deposited during the Late Triassic (mid to late Norian stage),Wellnhofer, P. (1991). "Summary of Triassic Pterosaurs." ''The Illustrated Encyclopedia of Pterosaurs.'' London, UK: Salamander Books Limited. p. 67. . making ''Eudimorphodon'' one of the oldest pterosaurs known.Cranfield, I. ''The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures''. London: Salamander Books, Ltd. Pp. 280–281. It had a wingspan of about and at the end of its long bony tail may have been a diamond-shaped flap like in the later '' Rhamphorhynchus''. If so, the flap may have helped it steer while maneuvering in the air. ''Eudimorphodon'' is known from several skeletons, including juvenile specimens. Discovery and species ''Eudimorphodon'' currently includes one species, the type ...
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Bergamodactylus
''Bergamodactylus'' is a putative genus of basal pterosaur which lived during the Late Triassic (early Norian) in the area of present-day Bergamo province in Italy. Its only species is ''Bergamodactylus wildi''. It was named in 2015 based on a pterosaur specimen which had previously been regarded as a juvenile ''Eudimorphodon'' or as identical to ''Carniadactylus''. Some Triassic pterosaur specialists consider the distinguishing features of ''Bergamodactylus'' to be invalid or insufficient to distinguish it from ''Carniadactylus'', and thus retain the specimen in that genus. History In 1978, Rupert Wild described a small pterosaur specimen in the collection of the ''Museo di Paleontologia dell´Università di Milano'', found near Cene, Lombardy. He referred to it as the "Milan Exemplar" and identified it as a juvenile of ''Eudimorphodon, Eudimorphodon ranzii''. Wild noted considerable differences with the latter's type specimen but these were explained as reflecting the young ...
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Rhamphorhynchus Muensteri
''Rhamphorhynchus'' (, from Ancient Greek ''rhamphos'' meaning "beak" and ''rhynchus'' meaning "snout") is a genus of long-tailed pterosaurs in the Jurassic period. Less specialized than contemporary, short-tailed pterodactyloid pterosaurs such as ''Pterodactylus'', it had a long tail, stiffened with ligaments, which ended in a characteristic soft-tissue tail vane. The mouth of ''Rhamphorhynchus'' housed needle-like teeth, which were angled forward, with a curved, sharp, beak-like tip lacking teeth, indicating a diet mainly of fish; indeed, fish and cephalopod remains are frequently found in ''Rhamphorhynchus'' abdominal contents, as well as in their coprolites. Although fragmentary fossil remains possibly belonging to ''Rhamphorhynchus'' have been found in England, Tanzania, and Spain, the best preserved specimens come from the Solnhofen limestone of Bavaria, Germany. Many of these fossils preserve not only the bones but impressions of soft tissues, such as wing membranes. Scatt ...
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Allkaruen
''Allkaruen'' (meaning "ancient brain") is a genus of "rhamphorhynchoid" pterosaur from the Early Jurassic Cañadon Asfalto Formation in Argentina. It contains a single species, ''A. koi.'' Description As demonstrated by a CT scan, the braincase of ''Allkaruen'' exhibits a unique set of traits that are intermediate between more basal pterosaurs such as ''Rhamphorhynchus'' and more derived pterodactyloid pterosaurs such as ''Anhanguera''. These traits are the position of the anterior semicircular canal of the inner ear, the relative orientations of the occiput and occipital condyle, the relative positions of the lateral margins of the flocculus and cerebral hemispheres, and the ratio between the length of the brain and the height of the hindbrain. In addition, the relative orientations of the frontal bone and lateral semicircular canal are more similar to ''Rhamphorhynchus'' than ''Anhanguera'', while the optic lobes are positioned lower than the forebrain as in pterodact ...
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Synapomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis ...
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Latin
Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the Roman Republic it became the dominant language in the Italian region and subsequently throughout the Roman Empire. Even after the fall of Western Rome, Latin remained the common language of international communication, science, scholarship and academia in Europe until well into the 18th century, when other regional vernaculars (including its own descendants, the Romance languages) supplanted it in common academic and political usage, and it eventually became a dead language in the modern linguistic definition. Latin is a highly inflected language, with three distinct genders (masculine, feminine, and neuter), six or seven noun cases (nominative, accusative, genitive, dative, ablative, and vocative), five declensions, four verb conjug ...
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