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Neoceratopsians
Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassic. The earliest known ceratopsian, '' Yinlong downsi'', lived between 161.2 and 155.7 million years ago.Holtz, Thomas R. Jr. (2011) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,'Winter 2010 Appendix./ref> The last ceratopsian species, ''Triceratops prorsus'', became extinct during the Cretaceous–Paleogene extinction event, . ''Triceratops'' is by far the best-known ceratopsian to the general public. It is traditional for ceratopsian genus names to end in "''-ceratops''", although this is not always the case. One of the first named genera was '' Ceratops'' itself, which lent its name to the group, although it is considered a ''nomen dubium'' today as its fossil remains have no distinguishing charac ...
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Microceratus
''Microceratus'' (meaning "small-horned") is a genus of small ceratopsian dinosaur that lived in the Cretaceous period in Asia. It walked on two legs, had short front arms, a characteristic ceratopsian frill and beak-like mouth, and was around long. It was one of the first ceratopsians, or horned dinosaurs, along with '' Psittacosaurus'' in Mongolia. Discovery The type species, ''Microceratops gobiensis'', was first described by Bohlin in 1953, and so was the second species, ''M. sulcidens'', which may belong to ''Asiaceratops'' instead. However, the generic name was already preoccupied by an ichneumon wasp (subfamily Cryptinae) with the same name. Though much of the material has since been reassigned to the genus '' Graciliceratops'', a replacement name ''Microceratus'' was created by Mateus in 2008 for the type specimen. Classification ''Microceratus'' belonged to the Ceratopsia (Ancient Greek for "horned face"), a group of herbivorous dinosaurs with parrot-like beaks w ...
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Archaeoceratops
''Archaeoceratops'', meaning "ancient horned face", is a genus of basal neoceratopsian dinosaur from the Early Cretaceous (Aptian stage) of north central China. It appears to have been bipedal and quite small, reaching in length and in body mass. It had a comparatively large head but no horns, possessing only a small bony frill projecting from the back of its head. Discovery and species Two specimens were found in the Middle Gray Unit of the Xinminbao Group, Gongpoquan Basin of the Mazong Shan area of Gansu Province, north central China. The type species, ''A. oshimai'', was named by Dong Zhiming and Azuma in 1997. It is the first basal neoceratopsian discovered in this area. The type specimen, IVPP V11114, consists of a partially complete skeleton including skull, caudal vertebrae, pelvis, and most of a hind foot. The second specimen (paratype), IVPP V11115, consists of an incomplete skeleton with a relatively well preserved caudal series, a partial hind limb, and a com ...
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Koreaceratops
''Koreaceratops'' () is a genus of basal ceratopsian dinosaur discovered in Albian-age Lower Cretaceous rocks of South Korea. Discovery It is based on KIGAM VP 200801, an articulated series of 36 caudal vertebrae associated with partial hind limbs and ischia. This specimen was found in a sandstone block that had been incorporated into the Tando dam at Hwaseong City; the way the specimen is cut off suggests that more of it was present before quarrying. The dam was built in 1994, and the bones were first brought to the attention of paleontologists in 2008, after a public official noticed them. The specimen came from the Sihwa Formation. ''Koreaceratops'' was described by Yuong-Nam Lee and colleagues in 2011. The genus name is a combination of "Korea" and the Greek ''κέρας'' (''keras'') meaning 'horn' and ''ὄψις'' (''opsis'') meaning 'face'. The type species is ''K. hwaseongensis'', named after Hwaseong City. Classification ''Koreaceratops'' is notable fo ...
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Beg Tse
''Beg'' (after the Himalayan war deity Beg-tse) is a genus of neoceratopsian dinosaur from the early Cretaceous period of Mongolia. The genus contains a single species, ''Beg tse'', known from a partial skull and very fragmentary postcrania. ''Beg'' represents the most basal neoceratopsian currently known. Discovery and naming The holotype, IGM 100/3652, was discovered in 2015 near the town of Tsogt-Ovoo in the Ömnögovi Province of Mongolia. Described from the Ulaanoosh Formation, the specimen is dated to 113 to 94 million years ago, at the boundary of the Lower and Upper Cretaceous. ''Beg'' is named after Beg-tse, a Himalayan deity who is the god of war in Mongolian culture. The deity is often depicted with a rugose face and/or body, similar to the appearance of the preserved skull of the dinosaur. Description Based on the size of the skull, about long, ''Beg'' was most likely a medium-sized basal ceratopsian, similar in size to '' Yinlong'' and '' Liaoceratop ...
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Liaoceratops
''Liaoceratops'', meaning "Liaoning horned face", is a ceratopsian dinosaur believed to be an early relative of the horned ceratopsids. It lived in the Early Cretaceous, 126 million years ago. It was discovered in China by a team of American and Chinese scientists. ''Liaoceratops'' was much smaller than its later relatives, but offers a glimpse into the early evolution of this group of dinosaurs. Discoveries and naming ''Liaoceratops'' was discovered in the famous Liaoning Province of China, where several fossils of feathered dinosaurs have also been collected. The type species ''Liaoceratops yanzigouensis'' was in 2002 named and described by Xu Xing, Peter Makovicky, Wang Xiaolin, Mark Norell and You Hailu. The generic name is derived from Liaoning and the Greek ''keras'', "horn" and ''ops'', "face". The specific name refers to the town Yanzigou. The holotype IVPP V12738 has been found in the Yixian Formation dating from the Barremian. These beds have also yielded fossil ...
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Mosaiceratops
''Mosaiceratops'' is a genus of ceratopsian, described by Zheng, Jin & Xu in 2015 and found in the Xiaguan Formation of Neixiang County. ''Mosaiceratops'' lived in the upper Cretaceous in what is now the Henan Province of China. Although phylogenetic analyses have found ''Mosaiceratops'' to be the most basal neoceratopsian, the authors noted that several features in the premaxilla and nasal bones are shared with '' Psittacosaurus'', indicating that neoceratopsians evolved premaxillary teeth twice and that ''Psittacosaurus'' is not as primitive as previously thought.Zheng, W., Jin, X., & Xu, X. (2015)A psittacosaurid-like basal neoceratopsian from the Upper Cretaceous of central China and its implications for basal ceratopsian evolution ''Scientific reports'', 5. article number 14190: 1-9; doi:10.1038/srep14190 Discovery On the westbank of the river Tuanhe in Neixiang in Henan the skeleton was discovered of a small ceratopian. The fossil was prepared by Sheng Yiming and Yu Ch ...
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Notoceratops
''Notoceratops'' (meaning "southern horned face") is a nomen dubium, dubious genus of extinct ornithischian dinosaur based on an incomplete, toothless left dentary (now lost) from the Late Cretaceous of Patagonia (in Argentina), probably dating to the Campanian or Maastrichtian. It was most likely a ceratopsian and it was found in the Lago Colhué Huapí Formation, Lago Colhué Huapi Formation. Discovery and naming In 1918, palaeontologist Augusto Tapia (1893–1966) discovered the genus holotype. He also named the type species, ''N. bonarellii'' (originally spelt as ''Notoceratops Bonarelli''), in 1918. The generic name is derived from Greek ''notos'', "the south", ''keras'', "horn" and ''ops'', "face". The specific name (zoology), specific name honours Guido Bonarelli (1871-1951), who advised Tapia in his study of the find. By present conventions the epithet is spelled ''bonarellii'', thus without a capital B. In many later publications the specific name is misspelled "bonarelli ...
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Serendipaceratops
''Serendipaceratops'' (meaning "serendipitous horned face") is a genus of herbivorous ornithischian dinosaur, possibly an ankylosaur, from the early Cretaceous Period of Australia. The type species, ''S. arthurcclarkei'', was named in 2003. Discovery and species The first known bone from ''Serendipaceratops'', an ulna (lower arm bone), was in 1993 discovered in Australia near Kilcunda, on the south-east coast of Victoria. The find took place during the " Dinosaur Cove" project, uncovering remains at the basis of a cliff face at the shoreline; the fossil was excavated in the "Arch", a small area of sea-floor protected from the waves by a dam. In 2003, the type species ''Serendipaceratops arthurcclarkei'' was named and described by Tom Rich and Patricia Vickers-Rich.T. Rich and P. Vickers-Rich. 2003. Protoceratopsian? ulnae from Australia. ''Records of the Queen Victoria Museum Launceston'' 113:1-12 Initially, the discoverers had not considered the possibility that the fossil ...
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Late Jurassic
The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name "Malm" indicates rocks of Late Jurassic age. In the past, ''Malm'' was also used to indicate the unit of geological time, but this usage is now discouraged to make a clear distinction between lithostratigraphic and geochronologic/chronostratigraphic units. Subdivisions The Late Jurassic is divided into three ages, which correspond with the three (faunal) stages of Upper Jurassic rock: Paleogeography During the Late Jurassic Epoch, Pangaea broke up into two supercontinents, Laurasia to the north, and Gondwana to the south. The result of this break-up was the spawning of the Atlantic Ocean. However, at this time, the Atlantic Ocean was relatively narrow. Life forms of the epoch This epoch is well known for many famous types of dinosau ...
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Yamaceratops
''Yamaceratops'' is a genus of primitive ceratopsian that lived in Asia during the Late Cretaceous period in what is now the Javkhlant Formation. Initially, the rocks where it was found in were thought to be from the Early Cretaceous, but the age was reevaluated in 2009. It was a relatively small dinosaur, reaching in length and in body mass. The type species, ''Yamaceratops dorngobiensis'', was described by P. J. Makovicky and M. A. Norell in September, 2006. The authors consider the animal to have had an intermediate phylogenetic position between ''Liaoceratops'' and ''Archaeoceratops'' within Neoceratopia. Examination of the frill of ''Yamaceratops'' has convinced the authors that the frill was not used for display, and that the fossils " intat a more complex evolutionary history for ceratopsian frills." The genus name refers to Yama, a Tibetan Buddhist deity; the species name to the Eastern Gobi. The holotype IGM 100/1315 consists of a partial skull; other material has ...
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Leptoceratopsidae
Leptoceratopsidae is an extinct family of neoceratopsian dinosaurs from Asia, North America and Europe. Leptoceratopsids resembled, and were closely related to, other neoceratopsians, such as the families Protoceratopsidae and Ceratopsidae, but they were more primitive and generally smaller. Phylogeny Leptoceratopsidae was originally named by Franz Nopcsa von Felső-Szilvás in 1923 as a subfamily Leptoceratopsinae, and its type species is '' Leptoceratops gracilis''. Mackovicky, in 2001, defined it as a stem-based taxon and a family consisting of ''Leptoceratops gracilis'' and all species closer to ''Leptoceratops'' than to '' Triceratops horridus''.Makovicky, P.J. 2001. A ''Montanoceratops cerorhynchus'' (Dinosauria: Ceratopsia) braincase from the Horseshoe Canyon Formation of Alberta, In: Tanke, D.H. & Carpenter, K. (Eds.). ''Mesozoic Vertebrate Life''. Bloomington: Indiana University Press. Pp. 243-262. The cladogram below follows the topologies from a 2015 analysis b ...
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