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Meridiolestida
Meridiolestida is an extinct clade of mammals known from the Cretaceous and Cenozoic of South America and possibly Antarctica. They represented the dominant group of mammals in South America during the Late Cretaceous. Meridiolestidans were morphologically diverse, containing both small insectivores such as the "sabretooth-squirrel" '' Cronopio,supplementary information
'' as well as the clade Mesungulatoidea/Mesungulatomorpha, which ranged in size from the shrew-sized '''' to the dog-sized '' Peligrotherium.'' ...
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Dryolestoidea
Dryolestida is an extinct order of mammals, primarily and possibly exclusively known from the Jurassic and Cretaceous. They are considered members of the clade Cladotheria, close to the ancestry of therian mammals. It is also believed that they developed a fully mammalian jaw and also had the three middle ear bones. Most members of the group, as with most Mesozoic mammals, are only known from fragmentary tooth and jaw remains. The taxonomic composition of the group is contested. Aside from the uncontroversial Dryolestidae and the possibly paraphyletic Paurodontidae, which were small insectivores, known from the Middle Jurassic to Early Cretaceous of Laurasia, the Meridiolestida, a diverse group of mammals including both small insectivores and mid-large sized herbivores known from the Late Cretaceous to Miocene of South America and possibly Antarctica, are also often included. However, in many phylogenetic analyses, Meridiolestida are recovered as an unrelated group of clado ...
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Necrolestes
''Necrolestes'' ("grave robber" or "thief of the dead") is an extinct genus of mammals, which lived during the Early Miocene in what is now Argentine Patagonia. It is the most recent known genus of Meridiolestida, an extinct group of mammals more closely related to therians (marsupials and placentals) than to monotremes that dominated South America during the Late Cretaceous. It contains two species, ''N. patagonensis'' and ''N. mirabilis,'' The type species ''N. patagonensis'' was named by Florentino Ameghino in 1891 based on remains found by his brother, Carlos Ameghino in Patagonia. Fossils of ''Necrolestes'' have been found in the Sarmiento and Santa Cruz Formations.''Necrolestes''
at Fossilworks.org
Its morph ...
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Cronopio (mammal)
''Cronopio'' is an extinct genus of small insectivorous mammal known from the early Late Cretaceous of the Río Negro region in Argentina. Its only species is ''Cronopio dentiacutus''.supplementary information
It belongs to the Meridiolestida, an extinct group of mammals widespread in South America during the Late Cretaceous, which are more closely related to modern s and
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Cladotheria
Cladotheria is a clade (sometimes ranked as a legion) of mammals. It contains modern therian mammals (marsupials and placentals) and several extinct groups, such as the dryolestoids, amphitheriids and peramurids. The clade was named in 1975 by Malcolm McKenna. In 2002, it was defined as a node-based taxon containing "the common ancestor of dryolestids and living therians, plus all its descendants". A different, stem-based definition was given in 2013, in which Cladotheria contains all taxa that are closer to ''Mus musculus'' (the house mouse) than to the "symmetrodont" '' Spalacotherium tricuspidens''. Description Early cladotherians can be distinguished from other mammals by a number of derived traits (apomorphies). Their teeth differed from those of the "symmetrodonts" by the evolution of a talonid shelf ( hypoflexid) on the lower molars, which occluded with the paracone of the corresponding upper molars. A true talonid basin, allowing for the crushing and grinding of f ...
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Reigitherium
''Reigitherium'' was a mammal that lived during the Late Cretaceous, in the (Late Campanian-Maastrichtian). Its fossils have been found in the Los Alamitos and the La Colonia Formations of Argentina. Description The original specimen of ''Reigitherium'' was a fragmentary single molar tooth, with a lot of the surface detail damaged. It was mistakenly identified as an upper left molar, but new material - including a whole tooth row of this species - clarifies that it was a lower right tooth. ''Reigitherium'' was a small mammal with simple premolars that increased in size along the tooth row to an enlarged fourth premolar. The molar teeth them decreased in size along the tooth row. Taxonomy ''Reigitherium'' has proven difficult to classify until recently, because the original fossil material was sparse, damaged, and difficult to identify. It was initially thought to be a dryolestid mammal when described in 1990. Ten years later, Pascual et al. argued that it was a docodont ...
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Peligrotherium
''Peligrotherium'' is an extinct meridiolestidan, and the sole member of the family Peligrotheriidae, from the Paleocene of Patagonia, originally interpreted as a stem-ungulate (though it did co-exist with early meridiungulates). Its remains have been found in the Salamanca Formation.''Peligrotherium''
at Fossilworks.org
It was a -sized mammal, among the largest of all dryolestoids (as well as the largest South American mammal), and closely related to



Dryolestidae
Dryolestidae is an extinct family of Mesozoic mammals, known from the Middle Jurassic to the Early Cretaceous of the North Hemisphere. The oldest known member, '' Anthracolestes'', is known from the Middle Jurassic Itat Formation of Western Siberia, but most other representatives are known from the Late Jurassic of North America and the Late Jurassic and Early Cretaceous of Europe. Most members are only known from isolated teeth and jaw fragments. Like many other groups of early mammals, they are though to have been insectivores. They are generally classified in Cladotheria, meaning that they are considered to be more closely related to marsupials and placentals than to monotremes. They are placed as part of the broader Dryolestida, which also includes the (possibly paraphyletic) Paurodontidae, and also sometimes the South American-Antarctic Meridiolestida, which are often considered unrelated cladotherians. Dyolestidae is not based on a phylogenetic definition, but instead on t ...
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Miocene
The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. ...
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Antarctic Peninsula
The Antarctic Peninsula, known as O'Higgins Land in Chile and Tierra de San Martín in Argentina, and originally as Graham Land in the United Kingdom and the Palmer Peninsula in the United States, is the northernmost part of mainland Antarctica. The Antarctic Peninsula is part of the larger peninsula of West Antarctica, protruding from a line between Cape Adams ( Weddell Sea) and a point on the mainland south of the Eklund Islands. Beneath the ice sheet that covers it, the Antarctic Peninsula consists of a string of bedrock islands; these are separated by deep channels whose bottoms lie at depths considerably below current sea level. They are joined by a grounded ice sheet. Tierra del Fuego, the southernmost tip of South America, is about away across the Drake Passage. The Antarctic Peninsula is in area and 80% ice-covered. The marine ecosystem around the western continental shelf of the Antarctic Peninsula (WAP) has been subjected to rapid climate change. Over the pas ...
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Austrotriconodon
''Austrotriconodon'' is a mammal genus from the Campanian and Maastrichtian of South America. It currently contains only the type species, ''A. mckennai''. Originally assumed to be a eutriconodont, more recent studies have recovered it as a meridiolestidan dryolestoid. Description ''Austrotriconodon'' is known only from its teeth. Their similarity to the triconodont teeth of mammals such as eutriconodonts has caused taxonomic confusion and referral of this taxon to eutriconodonta; however, discovery of similar teeth in animals such as '' Cronopio'' has led to its reinterpretation as a meridolestidan. Discovery and species ''Austrotriconodon'' fossils were found in the Argentinean Los Alamitos Formation, dating to the Campanian or Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spann ...
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