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Macraucheniopsis
''Macraucheniopsis'' is an extinct genus of litoptern mammal belonging to the family Macraucheniidae from the Middle to Late Pleistocene of Argentina. It, along with ''Macrauchenia'', ''Neolicaphrium'', and ''Xenorhinotherium'' were among the youngest known genera of litopterns. Classification Cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ... showing the taxonomic relationships of ''Macraucheniopsis'': References {{Taxonbar, from=Q113946418 Macraucheniids Pleistocene mammals of South America Pleistocene Argentina Ensenadan Lujanian Fossils of Argentina Fossil taxa described in 1945 Prehistoric placental genera Quaternary Argentina ...
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Macraucheniinae
Macraucheniidae is a family in the extinct South American ungulate order Litopterna, that resembled camelids. They had three functional digits on the fore and hind feet, as well as elongate necks. The family is generally divided up into two subfamilies, Cramaucheniinae (which may be paraphyletic) and Macraucheniinae. The family shows retraction of the nasal region, most extremely to the top of the skull in derived macraucheniine taxa like ''Macrauchenia.'' which has been interpreted to have supported a probsocis, perhaps like that of a saiga antelope to filter dust, or a moose-like prehensile lip. The earliest unambiguous members of the family date to the late Oligocene around 30 million years ago. '' Polymorphis'' from the Eocene has historically been placed as a macraucheniid, but this has been doubted. Most early representatives had a body masses in the range of , though some like ''Llullataruca'' were as small as , and the last representatives of the family from the Pleistoc ...
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Macraucheniidae
Macraucheniidae is a family in the extinct South American ungulate order Litopterna, that resembled camelids. They had three functional digits on the fore and hind feet, as well as elongate necks. The family is generally divided up into two subfamilies, Cramaucheniinae (which may be paraphyletic) and Macraucheniinae. The family shows retraction of the nasal region, most extremely to the top of the skull in derived macraucheniine taxa like ''Macrauchenia.'' which has been interpreted to have supported a probsocis, perhaps like that of a saiga antelope to filter dust, or a moose-like prehensile lip. The earliest unambiguous members of the family date to the late Oligocene The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ... around 30 million years ago. '' Polymorphis'' from the Eoce ...
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Pternoconius
''Pternoconius'' is an extinct genus of macraucheniid litoptern from the Late Oligocene and Early Miocene of Argentina. Fossils of this genus have been found in the Sarmiento Formation of Argentina. Etymology The genus name, ''Pternoconius'', is a near-anagram of the closely related genus ''Coniopternium''. The species name refers to the similarity of the Eocene genus '' Polymorphis''. Species ''Pternoconius tournoueri'' In 1985, fossils found in the Early Miocene Colhué Huapí Member of the Sarmiento Formation were assigned to the genus under the name ''Pternoconius tournoueri'', consisting of a nearly complete hemimandible. ''Pternoconius bondi'' In 2016 a new species of macraucheniid litoptern was described coming from the Bajada del Diablo locality in the Sarmiento Formation, consisting of the anterior portion of the skull with the maxillary region, some fragments of nasal bones, a small portion of the left zygomatic process, and most of the upper dentition (i.e., ...
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Cullinia
''Cullinia'' is an extinct genus of litoptern, an order of South American native ungulates that included horse-like and camel-like animals such as ''Macrauchenia''. It is only known from fragmentary remains. ''Cullinia levis'' is known from Chasicoan remains found in the Arroyo Chasicó Formation of Argentina, and remains from the Brazilian state of Acre and the Huayquerian Ituzaingó Formation have been assigned to ''Cullinia'' sp. . History and naming ''Cullinia'' was described in 1931 by Cabrera and Kraglievich, from MLP 29-IX-178, a holotype containing several fragmentary remains including a mandible and a metatarsal. In 1995, Bond and López add to the holotype other remains from the upper dentition. It was named from the Araucanian word "cullin", meaning "animal". Description ''Cullinia'' was a slender, small Macraucheniidae. It had proportionally larger metapodials than ''Theosodon'', and its first lower molar was absent. Classification Below is a phylogenetic tree ...
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Macrauchenia
''Macrauchenia'' ("long llama", based on the now-invalid llama genus, ''Auchenia'', from Greek "big neck") is an extinct genus of large ungulate native to South America from the Pliocene or Middle Pleistocene to the end of the Late Pleistocene. It is a member of the extinct order Litopterna, a group of South American native ungulates distinct from the two orders which contain all living ungulates which had been present in South America since the early Cenozoic, over 60 million years ago, prior to the arrival of living ungulates in South America around 2.5 million years ago as part of the Great American Interchange. The bodyform of ''Macrauchenia'' has been described as similar to a camel, being one of the largest-known litopterns, with an estimated body mass of around 1 tonne. The genus gives its name to its family, Macraucheniidae, which like ''Macrauchenia'' typically had long necks and three-toed feet, as well as a retracted nasal region, which in ''Macrauchenia'' manifests as t ...
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Promacrauchenia
''Promacrauchenia'' is an extinct genus of macraucheniids that lived during the Late Miocene to Late Pliocene epochs of what is now Argentina and Bolivia. It belongs to the subfamily Macraucheniinae, which also includes ''Huayqueriana'', ''Macrauchenia'', and ''Xenorhinotherium.'' Fossils of this genus have been found in the Ituzaingó, Andalhuala, and Cerro Azul Formation The Cerro Azul Formation (), also described as Epecuén Formation, is a formation (geology), geological formation of Late Miocene (Tortonian, or Huayquerian in the South American land mammal age, SALMA classification) age in the Colorado Basin, Arg ...s of Argentina. Classification The genus ''Promacrauchenia'' was first described by Florentino Ameghino in 1904, on the basis of fossils found in Patagonia in lower Pliocene deposits and which Ameghino himself, years earlier, had described as a species of ''Macrauchenia'', as ''M. antiqua'' . In addition to the type species, ''Promacrauchenia'' ''antiqua'', ...
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Xenorhinotherium
''Xenorhinotherium'' is an extinct genus of macraucheniine macraucheniids, native to northern South America during the Pleistocene and Holocene epoch, closely related to ''Macrauchenia'' of Patagonia. The type species is ''X. bahiense''.''Xenorhinotherium''
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Taxonomy

Some authors have proposed that the genus ''Xenorhinotherium'' a synonym of ''Macrauchenia'', though this has not been widely accepted. The name ''Xenorhinotherium'' means "Strange-Nosed Beast" and ''bahiense'' refers to the Brazilian state of

Cramauchenia
''Cramauchenia'' is an extinct genus of litoptern South American ungulate. ''Cramauchenia'' was named by Florentino Ameghino. The name has no literal translation. Instead, it is an anagram of the name of a related genus ''Macrauchenia''. This genus was initially discovered in the Sarmiento Formation in the Chubut Province, in Argentina, and later it was found in the Chichinales Formation in the Río Negro Province and the Cerro Bandera Formation in Neuquén, also in Argentina, in sediments assigned to the SALMA Colhuehuapian (in the Early Miocene), as well as the Agua de la Piedra Formation in Mendoza, in sediments dated to the Deseadan (during the Late Oligocene). In 1981 Soria made ''C. insolita'' a junior synonym of ''C. normalis''. A specimen of ''C. normalis'' was described in 2010 from Cabeza Blanca (Chubut, Argentina) in the Sarmiento Formation, in sediments assigned to the Deseadan SALMA (Upper Oligocene). Description This animal had an appearance vaguely similar to th ...
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Scalabrinitherium
''Scalabrinitherium'' is an extinct genus of mammals of the family Macraucheniidae. Fossils of this animal were found among the fossils of prehistoric xenarthrans in the Ituzaingó Formation of Argentina.''Scalabrinitherium''
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Description

This animal was rather similar to a with a slightly heavy build; the was long and low, the front teeth were slightly spatulate, and the nasal aperture set far back. It is possible tha ...
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Huayqueriana
''Huayqueriana'' is an extinct genus of South American litoptern, related to ''Macrauchenia'', and belonging to the same family, Macraucheniidae. It was formerly known as ''Macrauchenidia latidens'', described in 1939 by Cabrera, but redefined as ''Huayqueriana'' in 2016 based on the earlier name convention of Rovereto 1914.Forasiepi et al., 2016, p.11Cabrera (A.), 1939. "Sobre vertebrados fósiles del Plioceno del Adolfo Alsina". ''Revista Museo de La Plata'', II (n. Serie), Paleont.: 3–35. The genus is named after the Huayquerías Formation and the eponymous Huayquerian South American land mammal age defined at the formation. Classification Cladogram based in the Phylogenetics, phylogenetic analysis published by Schmidt ''et al''., 2014, showing the position of ''Huayqueriana'': References Bibliography

* Macraucheniids Miocene mammals of South America Huayquerian, Neogene Argentina Fossils of Argentina Cerro Azul Formation Fossil taxa described in 1914 Pre ...
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Theosodon
''Theosodon'' is an extinct genus of litoptern mammal from the Early to Middle Miocene of South America. Description ''Theosodon'' was long-legged with a long neck resembling modern llamas or guanacos. It was large for a litoptern, reaching up to in length and weighing up to . It had a long neck and tapir-like, three-toed feet, and like other litopterns and modern horses, tapirs and rhinos, it bore its weight on its middle toes. Extraordinarily, rather than having nostrils at the front of its head, ''Theosodon'' had its nostrils on the top of its snout, halfway between the forehead and the tip of the snout, and its nostrils pointed upwards rather than forwards, possibly as an adaptation for browsing on prickly vegetation. History and species ''Theosodon'' has been known since the 19th century, and by 1910 seven species had been described within the genus, all from the early Miocene Santa Cruz Formation in Argentina. The name ''Theosodon'' means "god tooth". The first f ...
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Paranauchenia
''Paranauchenia'' is an extinct genus of South American litopterns belonging to the family Macraucheniidae. It is known only from fossil finds in Argentina. It possessed three toes and long limbs. The species ''Paranauchenia denticulata'' lived in the Miocene epoch in Argentina.Ameghino, F. 1904a. Nuevas especies de mamíferos Cretáceos y Terciarios de la República Argentina. ''Anales de la Sociedad Científica Argentina'' 16: 193-208; 17: 162-175, 182-192, 225-240, 241-291. Fossils have been found in the Arroyo Chasicó and Ituzaingó Formations of Argentina.''Paranauchenia''
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Classification