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Limuloidea
Xiphosura (; , in reference to its sword-like telson) is an order of arthropods related to arachnids. They are more commonly known as horseshoe crabs (a name applied more specifically to the only extant family, Limulidae). They first appeared in the Hirnantian (Late Ordovician). Currently, there are only four living species. Xiphosura contains one suborder, Xiphosurida, and several stem-genera. The group has hardly changed in appearance in hundreds of millions of years; the modern horseshoe crabs look almost identical to prehistoric genera and are considered to be living fossils. The most notable difference between ancient and modern forms is that the abdominal segments in present species are fused into a single unit in adults. Xiphosura were historically placed in the class Merostomata, although this term was intended to encompass also the eurypterids, whence it denoted what is now thought to be an unnatural (paraphyletic) group (although this is a grouping recovered in some rece ...
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Horseshoe Crab
Horseshoe crabs are arthropods of the family Limulidae and the only surviving xiphosurans. Despite their name, they are not true crabs or even crustaceans; they are chelicerates, more closely related to arachnids like spiders, ticks, and scorpions. The body of a horseshoe crab is divided into three main parts: the cephalothorax, abdomen, and telson. The largest of these, the cephalothorax, houses most of the animal's eyes, limbs, and internal organs. It is also where the animal gets its name, as its shape somewhat resembles that of a horseshoe. Horseshoe crabs have been described as "living fossils", having changed little since they first appeared in the Triassic. Only four species of horseshoe crab are extant today. Most are marine, though the mangrove horseshoe crab is often found in brackish water, and the Atlantic horseshoe crab is resident in brackish estuarine ecosystems such as the Delaware and Chesapeake bays. Additionally, certain extinct species transitioned t ...
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Limulidae
Horseshoe crabs are arthropods of the family Limulidae and the only surviving xiphosurans. Despite their name, they are not true crabs or even crustaceans; they are chelicerates, more closely related to arachnids like spiders, ticks, and scorpions. The body of a horseshoe crab is divided into three main parts: the cephalothorax, abdomen, and telson. The largest of these, the cephalothorax, houses most of the animal's eyes, limbs, and internal organs. It is also where the animal gets its name, as its shape somewhat resembles that of a horseshoe. Horseshoe crabs have been described as "living fossils", having changed little since they first appeared in the Triassic. Only four species of horseshoe crab are extant today. Most are marine, though the mangrove horseshoe crab is often found in brackish water, and the Atlantic horseshoe crab is resident in brackish estuarine ecosystems such as the Delaware and Chesapeake bays. Additionally, certain extinct species transitioned to livi ...
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Hirnantian
The Hirnantian is the final internationally recognized stage of the Ordovician Period of the Paleozoic Era. It was of short duration, lasting about 2.1 million years, from to Ma (million years ago). The early part of the Hirnantian was characterized by cold temperatures, major glaciation, and a severe drop in sea level. In the latter part of the Hirnantian, temperatures rose, the glaciers melted, and sea level returned to the same or to a slightly higher level than it had been prior to the glaciation. Most scientists believe that this climatic oscillation caused the major extinction event that took place during this time. In fact, the Hirnantian (also known as the End Ordovician and the Ordovician-Silurian) mass extinction event represents the second largest such event in geologic history. Approximately 85% of marine (sea-dwelling) species died. Only the End-Permian mass extinction was larger. Unlike many smaller extinction events, however, the long-term consequences of t ...
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Paraphyletic
Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic grouping (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term received currency during the debates of the 1960s and 1970s accompanying the rise of cladistics, having been coined by zoologist Willi Hennig to apply to well-known taxa like Reptilia (reptiles), which is paraphyletic with respect to birds. Reptilia contains the last common ancestor of reptiles and all descendants of that ancestor exc ...
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Compound Eye
A compound eye is a Eye, visual organ found in arthropods such as insects and crustaceans. It may consist of thousands of ommatidium, ommatidia, which are tiny independent photoreception units that consist of a cornea, lens (anatomy), lens, and photoreceptor cells which distinguish brightness and color. The image perceived by this arthropod eye is a combination of inputs from the numerous ommatidia, which are oriented to point in slightly different directions. Compared with single-aperture eyes, compound eyes have poor image resolution; however, they possess a very large view angle and the ability to detect fast movement and, in some cases, the Polarization (waves), polarization of light. Because a compound eye is made up of a collection of ommatidia, each with its own lens, light will enter each ommatidium instead of using a single entrance point. The individual light receptors behind each lens are then turned on and off due to a series of changes in the light intensity during mov ...
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Labrum (arthropod Mouthpart)
The labrum is a flap-like structure that lies immediately in front of the mouth in almost all extant Euarthropoda. The most conspicuous exceptions are the Pycnogonida, which are probably chelicerates. In entomology, the labrum amounts to the "upper lip" of an insect mouth, the corresponding "lower lip" being the labium. The evolutionary origin, embryogenesis, and morphological development of the labrum have proved to be some of the most controversial and challenging topics in the study of arthropod head structures. Embryonic nature and origin of the labrum The labrum is innervated in crustaceans and insects from the tritocerebrum (the back of the brain). However, in development, its embryonic primordium often appears at the anterior of the head and migrates backwards towards its adult position. Furthermore, it often appears as a bilobed structure, with a set of muscles, nerves and gene expression in many ways similar to that of an appendage. This evidence has been used to ...
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Chelicera
The chelicerae () are the mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders. Commonly referred to as "jaws", chelicerae may be shaped as either articulated fangs, or as a type of pincers. Some chelicerae, such as those found on nearly all spiders, are hollow and contain (or are connected to) venom glands, used to inject venom into prey or a perceived threat. Both pseudoscorpions and harvestmen have additional structures on their chelicerae that are used for grooming (papillae in pseudoscorpions, cheliceral teeth in Opiliones). In '' Paratrechalea'', males and females have shown to have a chelicerae dimorphism, because the chelicerae is used as a mating signal for females. Types Chelicerae can be divided into three kinds: jackknife chelicerae, scissor chelicerae, and three-segmented chelate chelicerae. Jackknife chelicerae The jackknife chelicera is subchelate (with fixed finger much reduced or absent) an ...
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Carapace
A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the underside is called the plastron. In botany, a carapace refers to the hard outer cover of a seed which protects the inner embryo. Crustaceans In crustaceans, the carapace functions as a protective cover over the cephalothorax (i.e., the fused head and thorax, as distinct from the abdomen behind). Where it projects forward beyond the eyes, this projection is called a rostrum. The carapace is calcified to varying degrees in different crustaceans. Zooplankton within the phylum Crustacea also have a carapace. These include Cladocera, ostracods, and isopods, but isopods only have a developed "cephalic shield" carapace covering the head. Arachnids In arachnids, the carapace is formed by the fusion of prosomal tergites into a single pl ...
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Opisthosoma
The opisthosoma is the posterior part of the body in some arthropods, behind the prosoma ( cephalothorax). It is a distinctive feature of the subphylum Chelicerata (arachnids, horseshoe crabs and others). Although it is similar in most respects to an abdomen (and is often referred to as such), the opisthosoma is differentiated by its inclusion of the respiratory organs ( book lungs or book gills) and the heart. Segments The number of segments and appendages on the opisthosoma vary. Scorpions have 13, but the first is only seen during its embryological development. Other arachnids have fewer; harvestmen, for instance, have only ten. In general, appendages are absent or reduced, although in horseshoe crabs they persist as large plate-like limbs, called opercula or branchiophores, bearing the book gills, and that function in locomotion and gas exchange. In most chelicerates the opisthosomal limbs are greatly reduced and persist only as specialized structures, such as the silk- ...
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Prosoma
The cephalothorax, also called prosoma in some groups, is a tagma of various arthropods, comprising the head and the thorax fused together, as distinct from the abdomen behind. (The terms ''prosoma'' and ''opisthosoma'' are equivalent to ''cephalothorax'' and ''abdomen'' in some groups. The terms ''prosoma'' and ''opisthosoma'' may be preferred by some researchers in cases such as arachnids, where there is neither fossil nor embryonic evidence animals in this class have ever had separate heads and thoraxes, and where the ''opisthosoma'' contains organs atypical of a true ''abdomen'', such as a heart and respiratory organs.) The word ''cephalothorax'' is derived from the Greek words for head (, ') and thorax (, '). This fusion of the head and thorax is seen in chelicerates and crustaceans; in other groups, such as the Hexapoda (including insects), the head remains free of the thorax. In horseshoe crabs and many crustaceans, a hard shell called the carapace covers the cephalothorax. ...
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Paleozoic
The Paleozoic ( , , ; or Palaeozoic) Era is the first of three Era (geology), geological eras of the Phanerozoic Eon. Beginning 538.8 million years ago (Ma), it succeeds the Neoproterozoic (the last era of the Proterozoic Eon) and ends 251.9 Ma at the start of the Mesozoic Era. The Paleozoic is subdivided into six period (geology), geologic periods (from oldest to youngest), Cambrian, Ordovician, Silurian, Devonian, Carboniferous and Permian. Some geological timescales divide the Paleozoic informally into early and late sub-eras: the Early Paleozoic consisting of the Cambrian, Ordovician and Silurian; the Late Paleozoic consisting of the Devonian, Carboniferous and Permian. The name ''Paleozoic'' was first used by Adam Sedgwick (1785–1873) in 1838 to describe the Cambrian and Ordovician periods. It was redefined by John Phillips (geologist), John Phillips (1800–1874) in 1840 to cover the Cambrian to Permian periods. It is derived from the Ancient Greek, Greek ''palaiós'' (π� ...
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Ricinulei
Ricinulei is a small order of arachnids. Like most arachnids, they are predatory; eating small arthropods. They occur today in west-central Africa ('' Ricinoides'') and the Americas ('' Cryptocellus'' and '' Pseudocellus'') from Brazil to southern Texas, where they inhabit either leaf-litter or caves. As of 2022, 103 extant species of ricinuleids have been described worldwide, all in the single family Ricinoididae. Due to their obscurity they do not have a proper common-name, though in academic literature they are occasionally referred to as hooded tickspiders. In addition to the three living genera, Ricinulei has a fossil-record spanning over 300 million years, including fossils from the Late Carboniferous of Euramerica and the Cretaceous Burmese amber. Anatomy and physiology The most important general account of ricinuleid anatomy remains the 1904 monograph by Hans Jacob Hansen and William Sørensen. Useful further studies can be found in, e.g., the work of Pittard and Mitche ...
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