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Fluctuating Selection
Fluctuating selection is a mode of natural selection characterized by the fluctuation of the directional selection, direction of selection on a given phenotype over a relatively brief period of evolutionary time. For example, a species of plant may come in two varieties: one which prefers wetter soil and one which prefers dryer soil. During a period of wet years, the wet variety will be more fitness (biology), fit and produce more offspring, and thereby increase the frequency of wet-preferring plants. If this wet period is followed by drought, the dry variety will be selected for and its numbers will increase. As periods of dryness and wetness fluctuate, so too does selection on dry-preferring and wet-preferring plants. Fluctuating selection is also manifest at the genic level. Consider two alleles, A and B, which are found at the same locus (genetics), locus. Fluctuating selection dynamics are at play when selection favors A at time t0, B at t1 and A again at t2. Fluctuating select ...
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Ecological Niche
In ecology, a niche is the match of a species to a specific environmental condition. Three variants of ecological niche are described by It describes how an organism or population responds to the distribution of Resource (biology), resources and competitors (for example, by growing when resources are abundant, and when predators, parasites and pathogens are scarce) and how it in turn alters those same factors (for example, limiting access to resources by other organisms, acting as a food source for predators and a consumer of prey). "The type and number of variables comprising the dimensions of an environmental niche vary from one species to another [and] the relative importance of particular environmental variables for a species may vary according to the geographic and biotic contexts". See also Chapter 2: Concepts of niches, pp. 7 ''ff'' A Grinnellian niche is determined by the habitat in which a species lives and its accompanying Behavioral ecology, behavioral adaptations. ...
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Stabilizing Selection
Stabilizing selection (not to be confused with negative or purifying selection) is a type of natural selection in which the population mean stabilizes on a particular non-extreme trait value. This is thought to be the most common mechanism of action for natural selection because most traits do not appear to change drastically over time. Stabilizing selection commonly uses negative selection (a.k.a. purifying selection) to select against extreme values of the character. Stabilizing selection is the opposite of disruptive selection. Instead of favoring individuals with extreme phenotypes, it favors the intermediate variants. Stabilizing selection tends to remove the more severe phenotypes, resulting in the reproductive success of the norm or average phenotypes. This means that most common phenotype in the population is selected for and continues to dominate in future generations. History The Ukrainian and later Soviet evolutionary biologist Ivan Schmalhausen founded the theo ...
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Negative Selection (natural Selection)
In natural selection, negative selection or purifying selection is the selective removal of alleles that are deleterious. This can result in stabilising selection through the purging of deleterious genetic polymorphisms that arise through random mutations. Purging of deleterious alleles can be achieved on the population genetics level, with as little as a single point mutation being the unit of selection. In such a case, carriers of the harmful point mutation have fewer offspring each generation, reducing the frequency of the mutation in the gene pool. In the case of strong negative selection on a locus, the purging of deleterious variants will result in the occasional removal of linked variation, producing a decrease in the level of variation surrounding the locus under selection. The incidental purging of non-deleterious alleles due to such spatial proximity to deleterious alleles is called background selection. This effect increases with lower mutation rate but decreases ...
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Disruptive Selection
In evolutionary biology, disruptive selection, also called diversifying selection, describes changes in population genetics in which extreme values for a trait are favored over intermediate values. In this case, the variance of the trait increases and the population is divided into two distinct groups. In this more individuals acquire peripheral character value at both ends of the distribution curve. Overview Natural selection is known to be one of the most important biological processes behind evolution . There are many variations of traits, and some cause greater or lesser reproductive success of the individual. The effect of selection is to promote certain alleles, traits, and individuals that have a higher chance to survive and reproduce in their specific environment. Since the environment has a carrying capacity, nature acts on this mode of selection on individuals to let only the most fit offspring survive and reproduce to their full potential. The more advantageou ...
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Directional Selection
In population genetics, directional selection is a type of natural selection in which one extreme phenotype is favored over both the other extreme and moderate phenotypes. This genetic selection causes the allele frequency to shift toward the chosen extreme over time as allele ratios change from generation to generation. The advantageous extreme allele will increase in frequency among the population as a consequence of survival and reproduction differences among the different present phenotypes in the population. The allele fluctuations as a result of directional selection can be independent of the dominance of the allele, and in some cases if the allele is recessive, it can eventually become fixed in the population. Directional selection was first identified and described by naturalist Charles Darwin in his book ''On the Origin of Species'' published in 1859. He identified it as a type of natural selection along with stabilizing selection and disruptive selection. These types ...
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Frequency-dependent Selection
Frequency-dependent selection is an evolutionary process by which the fitness (biology), fitness of a phenotype or genotype depends on the phenotype or genotype composition of a given population. * In positive frequency-dependent selection, the fitness of a phenotype or genotype increases as it becomes more common. * In negative frequency-dependent selection, the fitness of a phenotype or genotype decreases as it becomes more common. This is an example of balancing selection. * More generally, frequency-dependent selection includes when biological interactions make an individual's fitness depend on the frequencies of other phenotypes or genotypes in the population. Frequency-dependent selection is usually the result of interactions between species (predation, parasitism, or competition), or between genotypes within species (usually competitive or symbiotic), and has been especially frequently discussed with relation to anti-predator adaptations. Frequency-dependent selection can lea ...
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Evolution Of Sexual Reproduction
Sexually reproducing animals, plants, fungi and protists are thought to have evolved from a common ancestor that was a single-celled eukaryotic species. Sexual reproduction is widespread in eukaryotes, though a few eukaryotic species have secondarily lost the ability to reproduce sexually, such as Bdelloidea, and some plants and animals routinely reproduce asexually (by apomixis and parthenogenesis) without entirely having lost sex. The evolution of sexual reproduction contains two related yet distinct themes: its ''origin'' and its ''maintenance.'' Bacteria and Archaea (prokaryotes) have processes that can transfer DNA from one cell to another (conjugation, transformation, and transduction), but it is unclear if these processes are evolutionarily related to sexual reproduction in Eukaryotes. In eukaryotes, true sexual reproduction by meiosis and cell fusion is thought to have arisen in the last eukaryotic common ancestor, possibly via several processes of varying suc ...
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Sexual Reproduction
Sexual reproduction is a type of reproduction that involves a complex life cycle in which a gamete ( haploid reproductive cells, such as a sperm or egg cell) with a single set of chromosomes combines with another gamete to produce a zygote that develops into an organism composed of cells with two sets of chromosomes ( diploid). This is typical in animals, though the number of chromosome sets and how that number changes in sexual reproduction varies, especially among plants, fungi, and other eukaryotes. In placental mammals, sperm cells exit the penis through the male urethra and enter the vagina during copulation, while egg cells enter the uterus through the oviduct. Other vertebrates of both sexes possess a cloaca for the release of sperm or egg cells. Sexual reproduction is the most common life cycle in multicellular eukaryotes, such as animals, fungi and plants. Sexual reproduction also occurs in some unicellular eukaryotes. Sexual reproduction does not occur in pro ...
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Ecosystem Model
An ecosystem model is an abstract, usually mathematical, representation of an ecological system (ranging in scale from an individual population, to an ecological community, or even an entire biome), which is studied to better understand the real system. Using data gathered from the field, ecological relationships—such as the relation of sunlight and water availability to photosynthetic rate, or that between predator and prey populations—are derived, and these are combined to form ecosystem models. These model systems are then studied in order to make predictions about the dynamics of the real system. Often, the study of inaccuracies in the model (when compared to empirical observations) will lead to the generation of hypotheses about possible ecological relations that are not yet known or well understood. Models enable researchers to simulate large-scale experiments that would be too costly or unethical to perform on a real ecosystem. They also enable the simulation of ec ...
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Evolutionary Arms Race
In evolutionary biology, an evolutionary arms race is an ongoing struggle between competing sets of co-evolving genes, phenotypic and behavioral traits that develop escalating adaptations and counter-adaptations against each other, resembling the geopolitical concept of an arms race. These are often described as examples of positive feedback.Dawkins, R. 1996. ''The Blind Watchmaker'' New York: W. W. Norton. Note: This book was also published by Penguin in 1991. While the text is identical, page numbers differ The co-evolving gene sets may be in different species, as in an evolutionary arms race between a predator species and its prey (Vermeij, 1987), or a parasite and its host. Alternatively, the arms race may be between members of the same species, as in the manipulation/sales resistance model of communication (Dawkins & Krebs, 1979) or as in runaway evolution or Red Queen effects. One example of an evolutionary arms race is in sexual conflict between the sexes, often describe ...
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Natural Selection
Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the Heredity, heritable traits characteristic of a population over generations. Charles Darwin popularised the term "natural selection", contrasting it with selective breeding, artificial selection, which is intentional, whereas natural selection is not. Genetic diversity, Variation of traits, both Genotype, genotypic and phenotypic, exists within all populations of organisms. However, some traits are more likely to facilitate survival and reproductive success. Thus, these traits are passed the next generation. These traits can also become more Allele frequency, common within a population if the environment that favours these traits remains fixed. If new traits become more favoured due to changes in a specific Ecological niche, niche, microevolution occurs. If new traits become more favoured due to changes in the ...
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