Eodiscina
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Eodiscina
Eodiscina is trilobite suborder. The Eodiscina first developed near the end of the Lower Cambrian period (late Atdabanian) and became extinct at the end of the Middle Cambrian. Species are tiny to small, and have a thorax of two or three segments. Eodiscina includes six families classified under one superfamily, Eodiscoidea. Taxonomy The Eodiscina are mostly considered the more primitive suborder of the Agnostida, and the Agnostina the more advanced. Some scholars do not consider the Agnostina true trilobites, and consequently rejected the idea that they were related to the Eodiscina. Consequently, these scientists have proposed to elevate the group to ordinal level, which would thus be called Eodiscida Kobayashi, 1939. Origin The oldest known eodiscoid is '' Tsunyidiscus''. The glabella of ''Tsunyidiscus'' is extremely similar to that of ''Dipharus clarki'', and distinct from all other eodiscoids. ''D. clarki'' is thought to represent an immature stage of the redlichioid '' ...
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Trilobite
Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period () and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized exoskeleton, they left an extensive fossil record. T ...
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Trilobita
Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period () and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized exoskeleton, they left an extensive fossil record. The ...
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Tsunyidiscus
''Tsunyidiscus'' is a trilobite belonging to the Suborder Eodiscina. ''Tsunyidiscus'' appeared near the end of the Lower Cambrian, during the late Atdabanian stage of geologic time and some collections suggest it may have survived into the Botomian. The genus is very small (up to 7mm), oculate and isopypous with a narrow dome-shaped glabella and a narrow bullet-shaped pygidial axis. Thorax consists of three segments. ''Tsunyidiscus'' is the only genus currently attributed to the family Tsunyidiscidae. Description Like other Agnostida the exoskeleton of ''Tsunyidiscus'' is diminutive and isopygous with 3 fulcrate thoracic segments. The cephalon has a strongly parabolic outline and maximum width (tr.) usually anterior to the genal angles. Glabella extremely narrow, lateral glabellar furrows usually obscure, with a rounded and expanded frontal glabellar lobe. The occipital ring (LO) is at least as long as L1, usually expanded laterally, and may bear a sharp posteriorly directed s ...
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Tsunyidiscidae
''Tsunyidiscus'' is a trilobite belonging to the Suborder Eodiscina. ''Tsunyidiscus'' appeared near the end of the Lower Cambrian, during the late Atdabanian stage of geologic time and some collections suggest it may have survived into the Botomian. The genus is very small (up to 7mm), oculate and isopypous with a narrow dome-shaped glabella and a narrow bullet-shaped pygidial axis. Thorax consists of three segments. ''Tsunyidiscus'' is the only genus currently attributed to the family Tsunyidiscidae. Description Like other Agnostida the exoskeleton of ''Tsunyidiscus'' is diminutive and isopygous with 3 fulcrate thoracic segments. The cephalon has a strongly parabolic outline and maximum width (tr.) usually anterior to the genal angles. Glabella extremely narrow, lateral glabellar furrows usually obscure, with a rounded and expanded frontal glabellar lobe. The occipital ring (LO) is at least as long as L1, usually expanded laterally, and may bear a sharp posteriorly directed s ...
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Weymouthiidae
The Weymouthiidae Kobayashi 1943 are an extinct family of eodiscinid agnostid trilobites. They lived during the late Lower Cambrian and earliest Middle Cambrian (Botomian to Delamaran) in the so-called ''Olenellus''- and ''Eokochaspis''-zones in the former paleocontinents of Laurentia, Avalonia, Gondwana. The Weymouthiidae are all blind and lack free cheeks. Taxonomy Ancestors The Weymouthiidae have developed from a stock within the Hebediscidae The Hebediscidae Kobayashi, 1944, are a family of trilobites belonging to the order Agnostida that lived during the Lower Cambrian (Atdabanian to Toyonian). They are small or very small, and have a thorax of two or three segments. The Hebediscida .... Descendants The Weymouthiidae are a paraphyletic family because the Agnostina suborder is nested within it, particularly in the clade that further consists of the genera ''Mallagnostus'', ''Chelediscus'', ''Tannudiscus'' and ''Jinghediscus''. The trend in the Weymouthiid ...
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Yukoniidae
Yukoniidae S. Zhang, 1980 'nom. transl''. et ''emend''. Jell, ''in'' Whittington ''et al''., 1997 ''ex'' Yukoniinae S. Zhang in W. Zhang, Lu ''et al''., 1980is a small family of trilobites, belonging to the Eodiscina. Type Genus and species ''Yukonia intermedia'' Palmer 1968 (Plate 2, figures 14, 17–19, 22, 23, 27, 28; text figure 4). Taxonomy The Yukoniidae probably descended from the Tsunydiscidae and gave rise to the Eodiscidae. cited in Description Yukoniidae are typically isopygous, belonging to the Superfamily Eodiscoidea. The narrow glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior t ... is usually parallel sided, anteriorly rounded and separated from smooth anterior border by broad (''sag''.) preglabellar field which occupies about 25% of cephalic length ...
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Agnostida
Agnostida is an order of arthropod which have classically been seen as a group of highly modified trilobites, though some recent research has doubted this placement. Regardless, they appear to be close relatives as part of the Artiopoda. They are present in the Lower Cambrian fossil record along with trilobites from the Redlichiida, Corynexochida, and Ptychopariida orders, and were highly diverse throughout the Cambrian. Agnostidan diversity severely declined during the Cambrian-Ordovician transition, and the last agnostidans went extinct in the Late Ordovician. Systematics The Agnostida are divided into two suborders — Agnostina and Eodiscina — which are then subdivided into a number of families. As a group, agnostids are isopygous, meaning their pygidium is similar in size and shape to their cephalon. Most agnostid species were eyeless. The systematic position of the order Agnostida within the class Trilobita remains uncertain, and there has been continuing de ...
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Pagetia Tajianensis Kaili 1
''Pagetia'' Walcott, 1916. is a small genus of trilobite, assigned to the Eodiscinid family Pagetiidae and which had global distribution during the Middle Cambrian. The genus contains 55 currently recognized species, each with limited spatial and temporal ranges. Type species ''Pagetia bootes'' Walcott, 1916, from the middle Cambrian (late Wuliuan or base Drumian) Burgess Shale fossil Lagerstätte. The species occurs with ''Ptychagnostus praecurrens'' and derives originally from the "greater phyllopod bed" of Walcott quarry on "Fossil Ridge" between Wapta Mountain and Mount Field, approximately 5 kilometres north of Field, British Columbia in the Canadian Rocky Mountains. The presence of ''Pt. praecurrens'', indicates correlation with the Swedish ''praecurrens'' Biozone (''Baltoparadoxides pinus'' Biosubzone, A2 of Westergård 1946, pp. 98–100) in the upper part of the ''oelandicus'' Superzone. Taxonomy Order: Agnostida Salter, 1864. Suborder: Eodiscina Kobayashi, ...
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Eodiscidae
Eodiscidae is a family of agnostid trilobites that lived during the final Lower Cambrian (late Toyonian) and the Middle Cambrian. They are small or very small, and have a thorax of two or three segments. Eodiscidae includes nine genera (see box). Taxonomy The Eodiscids probably descended from the agnostids of family Yukoniidae. cited in Description Like other agnostids, the body of eodiscids is diminutive, the headshield (or cephalon) and tailshield (or pygidium) are of approximately same size, with 2 or 3 thoracic segments in-between, each consisting of a horizontal inner portion that abruptly passes into an inclined outer portion (fulcrate). The central raised area of the cephalon (or glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior t ...) is narrow, usually with pa ...
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Calodiscidae
The Calodiscidae Kobayashi, 1943 'nom. transl.'' Öpik, 1975 ''ex'' Calodiscinae Kobayashi, 1943are a family of trilobites belonging to the order Agnostida that lived during the Lower Cambrian (Botomian and Toyonian). They are small or very small, and have a thorax of two or three segments. The Calodiscidae includes five genera (see box). Taxonomy The probable ancestors of the Calodiscidae are among the Tsunydiscidae. The Calodiscidae had no descendants. cited ''in'' Description Like all Agnostida, the Calodiscidae are diminutive and the headshield (or cephalon) and tailshield (or pygidium) are of approximately the same size (or isopygous) and outline. The central raised area of the cephalon (or glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior to ...) has parallel si ...
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Hebediscidae
The Hebediscidae Kobayashi, 1944, are a family of trilobites belonging to the order Agnostida that lived during the Lower Cambrian (Atdabanian to Toyonian). They are small or very small, and have a thorax of two or three segments. The Hebediscidae include five genera (see box). Taxonomy The Hebediscidae probably descended from the Tsunydiscidae and gave rise to the Weymouthiidae. cited in Description Like all Agnostida, the Hebediscidae are diminutive and the headshield (or cephalon) and tailshield (or pygidium) are of approximately the same size (or isopygous) and outline. In the Hebediscidae, the central raised area of the cephalon (or glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior to ...) is wide at its rear end, has parallel sides or tapers forward, and without t ...
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Peronopsidae
The Peronopsidae (which may also be called peronopsids) comprise the earliest family of the Agnostina suborder. Species of this family occurred on all paleocontinents. The earliest representatives of this family first occur just before the start of the Middle Cambrian, and the last disappeared just after the start of the Upper Cambrian. Distribution Peronopsidae are cosmopolitan. Temporal distribution Temporal distribution: * '' Archaeagnostus'', Toyonian ( ''Nephrolenellus multinodus''-zone) and Amgaian ('' Ovatoryctocara''-zone). * '' Eoagnostus'', Toyonian (''Nephrolenellus multinodus''-zone) and Amgaian (''Ovatoryctocara''-zone). * '' Peronopsis (Proacadagnostus)'', Amgaian (''Ovatoryctocara''- and lower ''Kounamkites''-zones). * '' Diplorrhina'', Amgaian (''Ovatoryctocara''- and ''Kounamkites''-zones) * '' Peronopsis (Peronopsis)'', Amgaian (''Ovatoryctocara''-, ''Kounamkites''- and ''Triplagnostus gibbus''-zones). * '' Pentagnostus (Pentagnostus)'', Amgaian (''Ovator ...
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