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Eodiscoidea
Eodiscina is trilobite suborder. The Eodiscina first developed near the end of the Lower Cambrian period (late Atdabanian) and became extinct at the end of the Middle Cambrian. Species are tiny to small, and have a thorax of two or three segments. Eodiscina includes six families classified under one superfamily, Eodiscoidea. Taxonomy The Eodiscina are mostly considered the more primitive suborder of the Agnostida, and the Agnostina the more advanced. Some scholars do not consider the Agnostina true trilobites, and consequently rejected the idea that they were related to the Eodiscina. Consequently, these scientists have proposed to elevate the group to ordinal level, which would thus be called Eodiscida Kobayashi, 1939. Origin The oldest known eodiscoid is '' Tsunyidiscus''. The glabella of ''Tsunyidiscus'' is extremely similar to that of ''Dipharus clarki'', and distinct from all other eodiscoids. ''D. clarki'' is thought to represent an immature stage of the redlichioid ''B ...
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Yukoniidae
Yukoniidae S. Zhang, 1980 'nom. transl''. et ''emend''. Jell, ''in'' Whittington ''et al''., 1997 ''ex'' Yukoniinae S. Zhang in W. Zhang, Lu ''et al''., 1980is a small family of trilobites, belonging to the Eodiscina. Type Genus and species ''Yukonia intermedia'' Palmer 1968 (Plate 2, figures 14, 17–19, 22, 23, 27, 28; text figure 4). Taxonomy The Yukoniidae probably descended from the Tsunydiscidae and gave rise to the Eodiscidae. cited in Description Yukoniidae are typically isopygous, belonging to the Superfamily Eodiscoidea. The narrow glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior to ... is usually parallel sided, anteriorly rounded and separated from smooth anterior border by broad (''sag''.) preglabellar field which occupies about 25% of cephalic length ...
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Agnostida
Agnostida is an order of arthropod which have classically been seen as a group of highly modified trilobites, though some recent research has doubted this placement. Regardless, they appear to be close relatives as part of the Artiopoda. They are present in the Lower Cambrian fossil record along with trilobites from the Redlichiida, Corynexochida, and Ptychopariida orders, and were highly diverse throughout the Cambrian. Agnostidan diversity severely declined during the Cambrian-Ordovician transition, and the last agnostidans went extinct in the Late Ordovician. Systematics The Agnostida are divided into two suborders — Agnostina and Eodiscina — which are then subdivided into a number of families. As a group, agnostids are isopygous, meaning their pygidium is similar in size and shape to their cephalon. Most agnostid species were eyeless. The systematic position of the order Agnostida within the class Trilobita remains uncertain, and there has been continuin ...
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Eodiscidae
Eodiscidae is a family of agnostid trilobites that lived during the final Lower Cambrian (late Toyonian) and the Middle Cambrian. They are small or very small, and have a thorax of two or three segments. Eodiscidae includes nine genera (see box). Taxonomy The Eodiscids probably descended from the agnostids of family Yukoniidae. cited in Description Like other agnostids, the body of eodiscids is diminutive, the headshield (or cephalon) and tailshield (or pygidium) are of approximately same size, with 2 or 3 thoracic segments in-between, each consisting of a horizontal inner portion that abruptly passes into an inclined outer portion (fulcrate). The central raised area of the cephalon (or glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior to ...) is narrow, usually with par ...
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Calodiscidae
The Calodiscidae Kobayashi, 1943 'nom. transl.'' Öpik, 1975 ''ex'' Calodiscinae Kobayashi, 1943are a family of trilobites belonging to the order Agnostida that lived during the Lower Cambrian (Botomian and Toyonian). They are small or very small, and have a thorax of two or three segments. The Calodiscidae includes five genera (see box). Taxonomy The probable ancestors of the Calodiscidae are among the Tsunydiscidae. The Calodiscidae had no descendants. cited ''in'' Description Like all Agnostida, the Calodiscidae are diminutive and the headshield (or cephalon) and tailshield (or pygidium) are of approximately the same size (or isopygous) and outline. The central raised area of the cephalon (or glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior to ...) has parallel sid ...
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Hebediscidae
The Hebediscidae Kobayashi, 1944, are a family of trilobites belonging to the order Agnostida that lived during the Lower Cambrian (Atdabanian to Toyonian). They are small or very small, and have a thorax of two or three segments. The Hebediscidae include five genera (see box). Taxonomy The Hebediscidae probably descended from the Tsunydiscidae and gave rise to the Weymouthiidae. cited in Description Like all Agnostida, the Hebediscidae are diminutive and the headshield (or cephalon) and tailshield (or pygidium) are of approximately the same size (or isopygous) and outline. In the Hebediscidae, the central raised area of the cephalon (or glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior to ...) is wide at its rear end, has parallel sides or tapers forward, and without tr ...
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Weymouthiidae
The Weymouthiidae Kobayashi 1943 are an extinct family of eodiscinid agnostid trilobites. They lived during the late Lower Cambrian and earliest Middle Cambrian (Botomian to Delamaran) in the so-called ''Olenellus''- and ''Eokochaspis''-zones in the former paleocontinents of Laurentia, Avalonia, Gondwana. The Weymouthiidae are all blind and lack free cheeks. Taxonomy Ancestors The Weymouthiidae have developed from a stock within the Hebediscidae. Descendants The Weymouthiidae are a paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ... family because the Agnostina suborder is nested within it, particularly in the clade that further consists of the genera ''Mallagnostus'', ''Chelediscus'', ''Tannudiscus'' and ''Jinghediscus''. The trend in the Weymouthiidae to ...
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Peronopsidae
The Peronopsidae (which may also be called peronopsids) comprise the earliest family of the Agnostina suborder. Species of this family occurred on all Supercontinent, paleocontinents. The earliest representatives of this family first occur just before the start of the Middle Cambrian, and the last disappeared just after the start of the Upper Cambrian. Distribution Peronopsidae are Cosmopolitan distribution, cosmopolitan. Temporal distribution Temporal distribution: * ''Archaeagnostus'', Toyonian (Trilobite zone#Subdivision of the Olenellus-zone, ''Nephrolenellus multinodus''-zone) and Amgan, Amgaian (''Trilobite zone, Ovatoryctocara''-zone). * ''Eoagnostus'', Toyonian (''Nephrolenellus multinodus''-zone) and Amgaian (''Ovatoryctocara''-zone). * ''Peronopsis, Peronopsis (Proacadagnostus)'', Amgaian (''Ovatoryctocara''- and lower ''Kounamkites''-zones). * ''Diplorrhina'', Amgaian (''Ovatoryctocara''- and ''Kounamkites''-zones) * ''Peronopsis, Peronopsis (Peronopsis)'', Amgaian ...
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Archaeagnostus
''Archaeagnostus'' is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It lived from the terminal Toyonian Cambrian Stage 4 is the still unnamed fourth stage of the Cambrian and the upper stage of Cambrian Series 2. It follows Cambrian Stage 3 and lies below the Wuliuan. The lower boundary has not been formally defined by the International Commissi ... to early Amgaian. Species belonging to this genus have been found in Eastern North America, Newfoundland, the Henson Gletscher Formation in Greenland, the Nekekit and Molodo Rivers in Eastern Siberia, and in Guizhou, China. References Cambrian trilobites of North America Peronopsidae Agnostida genera Extinct animals of North America Cambrian trilobites of Asia Paleozoic life of Quebec {{agnostida-stub Cambrian genus extinctions ...
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Thoracocare
''Thoracocare'' is a minute to very small (1.7 – 3.6 mm long) trilobite, that lived during part of the Middle Cambrian in what are today the states of Idaho, Nevada and Utah. It is the only trilobite known with just two thorax segments outside most members of the Agnostida order. It can be distinguished from Agnostida by the very wide subquadrate glabella (about half the cephalon), parallel-side or widening forward in the largest specimen, with the full front side touching the border. Two species are known, one, ''T. idahoensis'', only from pygidia. Etymology ''Thoracocare'' is derived from the Greek θώραξ (thorax) meaning "breastplate" and ἀκαρής (akares) meaning "tiny". Distribution *''T. minuta'' is known from the Middle Cambrian of the United States (''Glossopleura''-zone, Spence Tongue Member, Lead Bell Shale/Spence Shale Formation, Idaho, 42.0° N, 112.0° W). *''T. idahoensis'' was collected from the Middle Cambrian of the United States (''Glosso ...
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Corynexochida
Corynexochida is an order of trilobite that lived from the Lower Cambrian to the Late Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, w .... Like many of the other trilobite orders, Corynexochida contains many species with widespread characteristics. The middle region of the cephalon (the glabella) is typically elongate, with the sides often spreading forward (pestle-shaped). Some species have glabellae that are ''effaced'', meaning they are smooth and show little detail. The glabellar furrows (when not effaced) typically have a splayed arrangement. In most species, the hind pair on either side of the cephalon become spines that point sharply backwards, and the spinose tips of the anterior pairs of thoracic segments tend to become more and more forward directed toward the p ...
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Raphiophoridae
Raphiophoridae is a family of small to average-sized trilobites that first occurred at the start of the Ordovician and became extinct at the end of the Middle Silurian. Anatomy All raphiophorids are blind, with headshields (or cephalons) that are triangular to subcircular, and many carry long, trailing genal spines, a forward-directed rapier-like spine on the central raised area (or glabella), or both, with the glabella often inflated and the natural fracture lines (or sutures) of the cephalon coinciding with its margin. The thorax typically has five to seven segments (except for the genera ''Taklamakania'', ''Pseudampyxina'', ''Nanshanaspis'', and ''Kongqiangheia'', which have only 3). As mentioned before, all raphiophorids are blind. Many, if not most, genera have no eyes whatsoever, though a few, such as '' Lehnertia'', have vestigial tubercles that correspond to the compound eyes of their ancestors. Distribution Raphiophoridae currently includes two officially recog ...
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