Enaliktidae
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Enaliktidae
Enaliktidae is an extinct family of elongate arthropods known from the Silurian and Devonian periods, containing two genera, '' Enalikter'' and '' Bundenbachiellus.'' The taxonomic position of the family is uncertain. In its original description it was attributed to the Megacheira, a group of arthropods otherwise known from the Cambrian and Ordovician periods, due to them possessing uniramous frontal appendages with whip-like extensions, similar to the great appendages of megacheirans belonging to the family Leanchoiliidae like ''Leanchoilia.'' However, their placement as megacheirans has been questioned, as they arguably lack any defining apomorphies In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to hav ... of that group, as whether the great appendages of megacheirans and the frontal app ...
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Enalikter
''Enalikter'' ("scourger of the sea") is an extinct arthropod described from the middle Silurian Herefordshire Lagerstätte at the England–Wales border in UK. This genus is known from only one species, ''E. aphson''. ''Enalikter'' is described as late-living example of Megacheira, "great-appendage arthropod". It subsequently suggested to be an annelid by other researchers, however subsequent studies rejected this interpretation. Its interpretation as megacheiran arthropod has been questioned in later studies. Morphology ''Enalikter'' is small arthropod with length just around long. Head shield is subrectangular and lacked eyes. Median, unpaired antenna and three pairs of appendages can be seen on head, and a first appendage had three tapering flagella. Twelve trunk segments lack tergites, and is flexible to bend at least 90 degrees. Trunk appendages are biramous. The egg-shaped telson had two pairs of blade-like processes. Classification ''Enalikter'' is probably related ...
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Bundenbachiellus
''Bundenbachiellus'' is an extinct genus of arthropod described from the Lower Devonian Hunsrück Slate of Germany. This genus is known from only one species, ''B. giganteus''. Alongside its possible relative '' Enalikter'' from Silurian, it is possible that genus is late-living example of Megacheira, "great-appendage arthropod". History ''Bundenbachiellus giganteus'' was described in 1929 with the name of ''Megadactylus giganteus'', with incomplete specimen BSPG 1929 II 34, however genus name ''Megadactylus'' was already used for synonym of dinosaur ''Anchisaurus'', its genus name was changed into ''Bundenbachiellus'' in 1930. Second species, ''B. minor'' was also described, however that species is later considered as misinterpretation of a trilobite. Due to holotype specimen was incomplete, preserved without anterior section, ''Bundenbachiellus'' was once treated as partial fossil of cheloniellid arthropod ''Cheloniellon calmani'', and taxon itself as synonym of ''Cheloniel ...
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Megacheira
Megacheira ("great hands", also historically great appendage arthropods) is an extinct class of predatory arthropods defined by their possession of spined "great appendages". Their taxonomic position is controversial, with studies either considering them stem-group euarthropods, or stem-group chelicerates. The homology of the great appendages to the cephalic appendages of other arthropods is also controversial. Uncontested members of the group were present in marine environments worldwide from the lower Cambrian to the upper Ordovician. Morphology Megacheirans are defined by their possession of uniramous "great appendages", which are their first pair of head appendages. The first one or two proximalmost segments/ podomeres are spineless (it has been argued that the supposed first of the two proximal podomeres is actually an arthrodial membrane), while the remaining 3–4 more distal podomeres each typically bear a single upward pointing spine attached towards the distal en ...
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Silurian
The Silurian ( ) is a geologic period and system spanning 23.5 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the third and shortest period of the Paleozoic Era, and the third of twelve periods of the Phanerozoic Eon. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods ( myriapods, arachnids and hexapods) ...
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Devonian
The Devonian ( ) is a period (geology), geologic period and system (stratigraphy), system of the Paleozoic era (geology), era during the Phanerozoic eon (geology), eon, spanning 60.3 million years from the end of the preceding Silurian period at million years ago (Megaannum, Ma), to the beginning of the succeeding Carboniferous period at Ma. It is the fourth period of both the Paleozoic and the Phanerozoic. It is named after Devon, South West England, where rocks from this period were first studied. The first significant evolutionary radiation of history of life#Colonization of land, life on land occurred during the Devonian, as free-spore, sporing land plants (pteridophytes) began to spread across dry land, forming extensive coal forests which covered the continents. By the middle of the Devonian, several groups of vascular plants had evolved leaf, leaves and true roots, and by the end of the period the first seed-bearing plants (Pteridospermatophyta, pteridospermatophyt ...
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Cambrian
The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding Ediacaran period 538.8 Ma (million years ago) to the beginning of the Ordovician Period 486.85 Ma. Most of the continents lay in the southern hemisphere surrounded by the vast Panthalassa Ocean. The assembly of Gondwana during the Ediacaran and early Cambrian led to the development of new convergent plate boundaries and continental-margin arc magmatism along its margins that helped drive up global temperatures. Laurentia lay across the equator, separated from Gondwana by the opening Iapetus Ocean. The Cambrian marked a profound change in life on Earth; prior to the Period, the majority of living organisms were small, unicellular and poorly preserved. Complex, multicellular organisms gradually became more common during the Ediacaran, but it was not until the Cambrian that fossil diversity seems to rapidly ...
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Ordovician
The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era, and the second of twelve periods of the Phanerozoic Eon (geology), Eon. The Ordovician spans 41.6 million years from the end of the Cambrian Period Megaannum, Ma (million years ago) to the start of the Silurian Period Ma. The Ordovician, named after the Celtic Britons, Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same Rock (geology), rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed Stratum, strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official per ...
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Uniramous
The arthropod leg is a form of jointed appendage of arthropods, usually used for walking. Many of the terms used for arthropod leg segments (called podomeres) are of Latin origin, and may be confused with terms for bones: ''coxa'' (meaning hip, : ''coxae''), ''trochanter'', ''femur'' (: ''femora''), ''tibia'' (: ''tibiae''), ''tarsus'' (: ''tarsi''), ''ischium'' (: ''ischia''), ''metatarsus'', ''carpus'', ''dactylus'' (meaning finger), ''patella'' (: ''patellae''). Homologies of leg segments between groups are difficult to prove and are the source of much argument. Some authors posit up to eleven segments per leg for the most recent common ancestor of extant arthropods but modern arthropods have eight or fewer. It has been argued that the ancestral leg need not have been so complex, and that other events, such as successive loss of function of a ''Hox''-gene, could result in parallel gains of leg segments. In arthropods, each of the leg segments articulates with the next segmen ...
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Leanchoilia
''Leanchoilia'' is a megacheiran marine arthropod known from Cambrian deposits of the Burgess Shale in Canada and the Chengjiang biota of China. Description ''L. superlata'' was about long and had long, whip-like flagellae extending from its great appendages. Its internal organs are occasionally preserved within the substrate in three dimensions. Their two pairs of eyes are protected and covered by their exterior head shields, with two eyes being located on each side. Species Seven species are tentatively accepted today: ''L. superlata'' (the type species), ''L. persephone'' and ''L. protogonia'' from the Burgess Shale, ''L. illecebrosa'' and ''L. obesa'' from the Chengjiang biota, 'L. robisoni'' from Kaili, and ''L.''? ''hanceyi'' from the Spence Shale. ''L. superlata'' and ''L. persephone'' may however be examples of sexual dimorphism. Distribution 55 specimens of ''Leanchoilia'' are known from the Greater Phyllopod bed, where they comprise 0.1% of the community. S ...
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Apomorphies
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Determining ap ...
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Homology (biology)
In biology, homology is similarity in anatomical structures or genes between organisms of different taxa due to shared ancestry, ''regardless'' of current functional differences. Evolutionary biology explains homologous structures as retained heredity from a common descent, common ancestor after having been subjected to adaptation (biology), adaptive modifications for different purposes as the result of natural selection. The term was first applied to biology in a non-evolutionary context by the anatomist Richard Owen in 1843. Homology was later explained by Charles Darwin's theory of evolution in 1859, but had been observed before this from Aristotle's biology onwards, and it was explicitly analysed by Pierre Belon in 1555. A common example of homologous structures is the forelimbs of vertebrates, where the bat wing development, wings of bats and origin of avian flight, birds, the arms of primates, the front flipper (anatomy), flippers of whales, and the forelegs of quadrupedalis ...
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Enigmatic Arthropod Taxa
Enigmatic is an adjective meaning "mysterious" or "puzzling". It may also refer to: * ''Enigmatic'' (album), a 1970 album by Czesław Niemen * '' Enigmatic: Calling'', a 2005 album by Norwegian progressive metal band Pagan's Mind * Enigmatic scale, musical scale used by Verdi and others * "The Enigmatic", a song by Joe Satriani on the album '' Not of This Earth'' See also * Enigmatic leaf turtle, a species of Asian leaf turtle * Enigmatic moray eel, a species found in the Pacific and Indian Oceans * ''Glaresis ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy areas wo ...'', a genus of beetles sometimes called "enigmatic scarab beetles" * Enigma (other) {{disambig ...
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