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Dobsonfly Corydalus Cornutus Larvae
Dobsonflies are a subfamily of insects, Corydalinae, part of the Megalopteran family Corydalidae. The larvae (commonly called hellgrammites) are aquatic, living in streams, and the adults are often found along streams as well. The nine genera of dobsonflies are distributed in the Americas, Asia, and South Africa. Etymology The origin of the word "dobsonfly" is unclear. John Henry Comstock used the term in reference to these insects in his 1897 book ''Insect Life'', but did not explain it. He also mentioned that anglers used the word "hellgrammite" for the aquatic larvae they used as bait, but the origin of this term is also unknown. Description Adult dobsonflies are some of the largest non-Lepidopteran insects of temperate zones such as the United States and Canada, with a wingspan of up to in some species of ''Corydalus''.Stange, Lionel. "Alderflies and Dobsonflies." Encyclopedia of Entomology. 2nd ed. New York: Springer Publishing, 2008. The Asian ''Acanthacorydalis fru ...
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Corydalus Cornutus
The eastern dobsonfly, ''Corydalus cornutus'', is a large insect in the Corydalidae family. It is found in eastern North America in regions with fast-flowing streams where its aquatic larvae develop. These are known as hellgrammites and are among the top invertebrate predators in the streams in which they live. They are used by anglers as bait. Distribution The eastern dobsonfly is found in most of eastern North America. It is usually found near the swift flowing, unpolluted streams in which its larvae develop. Common names The origin of the word "dobsonfly" is unclear. John Henry Comstock used the term in reference to this species in his 1897 book ''Insect Life'',Comstock, John Henry (1897). Insect Life. Cornell University Library. Online. but did not explain it. He also mentioned that anglers use the word "hellgrammite" for the aquatic larvae they used as bait, but the origin of this term is also unknown.Turpin, TDobsonflies Look Vicious.Purdue Agriculture News Columns. Purdu ...
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Temperate Zones
In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout the year and more distinct seasonal changes compared to tropical climates, where such variations are often small and usually only have precipitation changes. In temperate climates, not only do latitudinal positions influence temperature changes, but sea currents, prevailing wind direction, continentality (how large a landmass is) and altitude also shape temperate climates. The Köppen climate classification defines a climate as "temperate" C, when the mean temperature is above but below in the coldest month to account for the persistency of frost. However, other climate classifications set the minimum at . Zones and climates The north temperate zone extends from the Tropic of Cancer (approximately 23.5° north latitude) to the Arcti ...
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Clades
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species ( extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organ ...
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Corydalinae
Dobsonflies are a subfamily of insects, Corydalinae, part of the Megalopteran family Corydalidae. The larvae (commonly called hellgrammites) are aquatic, living in streams, and the adults are often found along streams as well. The nine genera of dobsonflies are distributed in the Americas, Asia, and South Africa. Etymology The origin of the word "dobsonfly" is unclear. John Henry Comstock used the term in reference to these insects in his 1897 book ''Insect Life'', but did not explain it. He also mentioned that anglers used the word "hellgrammite" for the aquatic larvae they used as bait, but the origin of this term is also unknown. Description Adult dobsonflies are some of the largest non-Lepidopteran insects of temperate zones such as the United States and Canada, with a wingspan of up to in some species of '' Corydalus''.Stange, Lionel. "Alderflies and Dobsonflies." Encyclopedia of Entomology. 2nd ed. New York: Springer Publishing, 2008. The Asian '' Acanthacorydalis f ...
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Antenna (biology)
Antennae ( antenna), sometimes referred to as "feelers", are paired appendages used for Sensory system, sensing in arthropods. Antennae are connected to the first one or two Segmentation (biology), segments of the arthropod head. They vary widely in form but are always made of one or more jointed segments. While they are typically sensory organs, the exact nature of what they sense and how they sense it is not the same in all groups. Functions may variously include sensing tactition, touch, air motion, heat, vibration (sound), and especially insect olfaction, smell or gustation, taste. Antennae are sometimes modified for other purposes, such as mating, brooding, swimming, and even anchoring the arthropod to a substrate (biology), substrate. Larval arthropods have antennae that differ from those of the adult. Many crustaceans, for example, have free-swimming larvae that use their antennae for swimming. Antennae can also locate other group members if the insect lives in a group, lik ...
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Spermatophores
A spermatophore or sperm ampulla is a capsule or mass containing spermatozoa created by males of various animal species, especially salamanders and arthropods, and transferred in entirety to the female's ovipore during reproduction. Spermatophores may additionally contain nourishment for the female, in which case it is called a nuptial gift, as in the instance of bush crickets. In the case of the toxic moth ''Utetheisa ornatrix'', the spermatophore includes sperm, nutrients, and pyrrolizidine alkaloids which prevent predation because it is poisonous to most organisms. However, in some species such as the Edith's checkerspot butterfly, the "gift" provides little nutrient value. The weight of the spermatophore transferred at mating has little effect on female reproductive output. Arthropods Spermatophores are the norm in arachnids and several soil arthropods. In various insects, such as bush crickets, the spermatophore is often surrounded by a proteinaceous spermatophyla ...
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Sclerite
A sclerite ( Greek , ', meaning " hard") is a hardened body part. In various branches of biology the term is applied to various structures, but not as a rule to vertebrate anatomical features such as bones and teeth. Instead it refers most commonly to the hardened parts of arthropod exoskeletons and the internal spicules of invertebrates such as certain sponges and soft corals. In paleontology, a scleritome is the complete set of sclerites of an organism, often all that is known from fossil invertebrates. Sclerites in combination Sclerites may occur practically isolated in an organism, such as the sting of a cone shell. Also, they can be more or less scattered, such as tufts of defensive sharp, mineralised bristles as in many marine Polychaetes. Or, they can occur as structured, but unconnected or loosely connected arrays, such as the mineral "teeth" in the radula of many Mollusca, the valves of Chitons, the beak of Cephalopod, or the articulated exoskeletons of Arthropoda. ...
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Courtship Display
A courtship display is a set of display behaviors in which an animal, usually a male, attempts to attract a mate; the mate exercises choice, so sexual selection acts on the display. These behaviors often include ritualized movement ("dances"), vocalizations, mechanical sound production, or displays of beauty, strength, or agonistic ability. Male display In some species, males will perform ritualized movements to attract females. The male six-plumed bird-of-paradise ( ''Parotia lawesii'') exemplifies male courtship display with its ritualized "ballerina dance" and unique occipital and breast feathers that serve to stimulate the female visual system. In ''Drosophila subobscura,'' male courtship display is seen through the male's intricate wing scissoring patterns and rapid sidestepping. These stimulations, along with many other factors, result in subsequent copulation or rejection. In other species, males may exhibit courtship displays that serve as both visual and auditory sti ...
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Secondary Sex Characteristics
Secondary sex characteristics are features that appear during puberty in humans, and at sexual maturity in other animals. These characteristics are particularly evident in the sexually dimorphic phenotypic traits that distinguish the sexes of a species, but unlike the sex organs (primary sex characteristics), are not directly part of the reproductive system. Secondary sex characteristics are believed to be the product of sexual selection for traits which display fitness, giving an organism an advantage over its rivals in courtship and in aggressive interactions. Secondary sex characteristics include, for example, the manes of male lions, the bright facial and rump coloration of male mandrills, and horns in many goats and antelopes. These characteristics are believed to be produced by a positive feedback loop known as the Fisherian runaway produced by the secondary characteristic in one sex and the desire for that characteristic in the other sex. Male birds and fish of many spe ...
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Sexual Selection
Sexual selection is a mode of natural selection in which members of one biological sex choose mates of the other sex to mate with (intersexual selection), and compete with members of the same sex for access to members of the opposite sex (intrasexual selection). These two forms of selection mean that some individuals have greater reproductive success than others within a population, for example because they are more attractive or prefer more attractive partners to produce offspring. Successful males benefit from frequent mating and monopolizing access to one or more fertile females. Females can maximise the return on the energy they invest in reproduction by selecting and mating with the best males. The concept was first articulated by Charles Darwin who wrote of a "second agency" other than natural selection, in which competition between mate candidates could lead to speciation. The theory was given a mathematical basis by Ronald Fisher in the early 20th century. Sexual ...
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Mandible (insect Mouthpart)
Insect mandibles are a pair of appendages near the insect's mouth, and the most anterior of the three pairs of oral appendages (the labrum is more anterior, but is a single fused structure). Their function is typically to grasp, crush, or cut the insect's food, or to defend against predators or rivals. Insect mandibles, which appear to be evolutionarily derived from legs, move in the horizontal plane unlike those of vertebrates, which appear to be derived from gill arches and move vertically. Grasshoppers, crickets, and other simple insects The mouthparts of orthopteran insects are often used as a basic example of mandibulate (chewing) mouthparts, and the mandibles themselves are likewise generalized in structure. They are large and hardened, shaped like pinchers, with cutting surfaces on the distal portion and chewing or grinding surfaces basally. They are usually lined with teeth and move sideways. Large pieces of leaves can therefore be cut and then pulverized near the mouth ...
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Insect Wings
Insect wings are adult outgrowths of the insect exoskeleton that enable insects to fly. They are found on the second and third thoracic segments (the mesothorax and metathorax), and the two pairs are often referred to as the forewings and hindwings, respectively, though a few insects lack hindwings, even rudiments. The wings are strengthened by a number of longitudinal veins, which often have cross-connections that form closed "cells" in the membrane (extreme examples include the dragonflies and lacewings). The patterns resulting from the fusion and cross-connection of the wing veins are often diagnostic for different evolutionary lineages and can be used for identification to the family or even genus level in many orders of insects. Physically, some insects move their flight muscles directly, others indirectly. In insects with direct flight, the wing muscles directly attach to the wing base, so that a small downward movement of the wing base lifts the wing itself upward. Those ...
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